1946] 
Carpenter — Carboniferous insects 
119 
Order Palaeodictyoptera 
Family Spilapteridae Handlirsch 
Spilapteridae Handlirsch, 1906, Die fossilen Insekten p. 101. 
The insects included in this family have distinct cross veins, which 
usually form definite rows; a true archedictyon is absent. The 
anterior margin of the wings (fore and hind) tends to be concave, 
often strongly so; the shapes of the fore and hind wings may be 
very similar (e.g., Homaloneura) but more often the hind wing 
has a broader cubital-anal region (e.g., Dunbaria) ; in such cases, 
although the cubitus and anal veins may have a few more branches 
than in the fore wing, the venation is essentially alike in the two 
wings. The radial sector has at least six terminal branches, both 
MA and MP have at least two branches, and CuA has several 
branches. The wings are commonly marked with bands or spots. 
The family is known from Upper Carboniferous strata to the Upper 
Permian. I consider the families Dunbariidae Handlirsch (1937), 
Doropteridae Zalessky (1946) and Neuburgiidae Rohdendorf 
(1961) to be inseparable from the Spilapteridae. 
Genus Hojnaloneura Brongniart 
Homaloneura Brongniart, 1885, Bull. Soc. Amis Sci. Natur. Rouen, 1885:50- 
68 ; 1893, Recherches pour servir a l’histoire des insectes fossiles des temps 
primaires, p. 316. 
Homaloneurina Handlirsch, 1906, Die fossilen Insekten, p. 46. 
Homaloneurites Handlirsch, 1906, ibid., p. 47. 
Hojnaloneura belongs to that series of genera of the Spilapteridae 
in which the subcosta etxends nearly to the apex of the wings and 
in which there is no anastomosis of MA with Rs (or R) or of 
CuA with MP (or M). It is related to Doropteron (Permian, 
USSR), Permiakovia Martynov (Permian, USSR), Neuburgia 
Martynov (U. Carb., USSR) and Dunbaria Tillyard (L. Permian, 
Kansas). Unfortunately, our knowledge of even the wings of most 
of these genera is unsatisfactory; Permiakovia is known only from 
the hind wing and Doropteron and N euburgia are known from the 
fore wings or parts of the fore wings. Some synonymy may eventu- 
ally be necessary when more is known about these genera, but 
Explanation of Plate 16 
Homaloneura dabasinskasi, n.sp. Fore and hind wings; original drawing 
of holotype by Dr. J. Kukalova. C, costa ( + ), Sc, subcosta ( — ), Rl, radius 
( + ), Rs, radial sector ( — ), MA, anterior media ( + ), MP, posterior 
media ( — ), CuA, anterior cubitus ( + ), CuP, posterior cubitus ( — ), 1A, 
first anal ( + ). 
