138 
Psyche 
[September 
Species placed 
in P oner a by 
Heer 1849 
ctffinis 
crassinervis 
croatica 
elongatula 
fuliginosa (with subspecies 
oeningensis and radoboj) 
globosa 
longaeva 
nitida 
ventrosa 
Species placed 
Current 
in Poneropsis 
assignment 
by Heer 1867 
affinis 
Formicidae 
_ _ 
generis 
Poneropsis 
— 
Poneropsis 
elongatula 
Formicidae 
fuliginosa 
generis 
Poneropsis 

Poneropsis 
— 
Poneropsis 
nitida 
Poneropsis 
— 
Poneropsis 
incertae 
incertae 
them are now considered to belong in the genus. Popov’s citations 
are important, however, as he used some of the names originally 
assigned to Ponera by Heer, thus firmly establishing them in modern 
systematic nomenclature. Those involved are croatica , crassinervis 
(incorrectly spelled as crassicornis ) , ventrosa , longaeva and globosa. 
All of the additional thirteen species described in Poneropsis in 
1867 appear to be satisfactorily placed, except elongata, anthracina , 
imhoff, and stygia in which the wings are too incompletely preserved 
to allow allocation — they should be considered “Formicidae incertae 
generis” . 
A further species, Ponera veneraria, was described by Heer in his 
Urwelt der Schweiz (1865). This species was later transferred to 
Poneropsis in the 1879, second edition of the same work. On the 
basis of Heer’s 1865 figure I concur with Handlirsch (1908) that 
this species is best placed as Formicidae incertae generis. The name 
was misspelled “ vernaria ” by Handlirsch. 
Fossil Ponera described by authors other than Heer. 
The following list, as far as I am aware, includes all ant fossils 
allocated to Ponera by authors other than Heer. This includes those 
which have since been placed elsewhere by previous authors, whose 
reassignments are discussed below with my own opinions on the 
proper placement of all the species listed here. The appropriate ref- 
erences may be obtained in the bibliography. 
