PSYCHE 
Vol. 71 
December, 1964 
No. 4 
REVIEW OF THE CALVERTIELLIDAE, WITH 
DESCRIPTION OF A NEW GENUS FROM 
PERMIAN STRATA OF MORAVIA 
( PALAEODICTYOPTERA) 
By Jarmila Kukalova* 
Charles University, Prague 
The extinct order Palaeodictyoptera now includes some twenty 
families, which appear to represent many lines of evolution radiating 
in different directions. Most of them resemble the Ephemeroptera, 
at least superficially, more than any other order of living insects but 
a few show some features highly suggestive of the Odonata and 
Protodonata. 
The most notable of the latter is the family Calvertiellidae, now 
represented by Calvertiella Tillyard from Lower Permian deposits 
near Elmo, Kansas, and by Moraviptera Kukalova from the very 
base of the Lower Permian strata in Moravia (Zbysov). Recently, 
some remarkably preserved fossils belonging to a new genus were 
found in Moravia at the Obora locality. Since both fore and hind 
wings are preserved, they add a great deal to our knowledge of this 
group of Palaeodictyoptera. The present paper consists of a review 
of the Calvertiellidae and the description of the new genus Moravia 
from Czechoslovakia. 
Calvertiella was originally (1925) placed by Tillyard in the 
family Protagrionidae within the order Protodonata, but this was 
before the Protodonata (as well as Odonata) were known to lack 
the veins MA and CuA. The family Calvertiellidae, established by 
Martynov in 1931, was transferred to the Palaeodictyoptera by 
^Current address: Biological Laboratories, Harvard University. 
This research has been aided by a National Science Foundation Grant, 
No. NSFGP- 2038. I am much indebted to Professor F. M. Carpenter, who 
made it possible for me to study the type of Calvertiella and who was very 
helpful in the preparation of this paper. 
153 
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