156 
Psyche 
[December 
Handlirsch in 1937, along with the Protagrionidae. Since then, both 
families have been treated as Palaeodictyoptera. 1 
order Palaeodictyoptera golden berg 
Family Calvertiellidae Martynov, 1931 
Calvertiellidae, Martynov, 1931, Bull. Acad. Sci. URSS, 1:146 [Protodonata 
(Meganisoptera)] ; Handlirsch, 1937, Ann. Naturh. Mus. Wien, 48:82 
[Palaeodictyoptera]; Martynov, 1938, Trav. Inst, paleont., 7(4) :41 
[Protodonata]; Carpenter, 1943, Bull. Geol. Soc. Amer., 54:536 
[Palaeodictyoptera]; Laurentiaux, 1953, Traite de paleont. :424 [Palaeo- 
dictyoptera]; Carpenter, 1954, Bull. Mus. Comp. Zool., 108:786 
[Palaeodictyoptera]; Kukalova. 1955, Sbornik Ustr. Ust. geol., 21:571 
[Palaeodictyoptera]; Rodendorf, 1962, Osnovy paleont. :52 [Palaeodic- 
tyoptera]. 
Diagnosis. The following diagnosis of the family is based on a 
study of the type of Calvertiella permiana Tillyard, 1925, as well as 
on the material from Moravia. 
Fore wing: a very short convex veinlet present at the basal part 
of the costal area; Sc terminating on Ri slightly beyond mid-wing; 
Rs arising before the end of the first third of wing length, with 3-4 
main branches; a cuticular thickening (ct) 2 between R and M, at 
about the level of the origin of Rs; stems of R and M independent 
at the wing base, but approaching each other near the point of 
origin of Rs; M forking near the level of the origin of Rs; MA 
unbranched, MP branched; stem of Cu independent, its branches 
simple and strongly curved towards posterior margin ; CuA diverging 
towards M near the level of origin of Rs and fusing with it for 
some distance, or connected with it by means of a short, convex con- 
necting vein; in its further course, CuA runs very near to the 
posterior branch of MP; area between CuA and CuP broad; 6 anal 
veins, at least the first 3 arising from one independent stem, all 
strongly arched, especially the proximal ones; cross veins more or 
less numerous, forming a reticulation in at least some areas of the 
wing; intercalary sectors more or less definitely indicated in the 
radial and medial areas. 
Hind wing: broad basally and triangular in shape ( Moravia , 
n.g.) ; anterior margin concave; basal veinlets in costal area some- 
what better developed than in the fore wing; Sc terminating much 
1 For a discussion of the nomenclatural problems involving the name 
Protodonata see Carpenter, 1943 and 1954. 
2 Not known in Moraviptera because of poor preservation in the region 
of the origin of Rs. 
