1964] 
Kukalova — Calvertiellidae 
157 
as in fore wing; Rs arising slightly more basally than in the fore 
wing, with about 4 main branches; cuticular thickening as in fore 
wing; stem of M independent, forked near the level of origin of 
Rs; MA unbranched, MP branched; stem of Cu independent, 
dividing into two simple branches, arched strongly towards posterior 
margin; CuA diverging towards M as in the fore wing, and then 
continuing close to the posterior branch of MP; 4 anal veins, at 
least 3 arising from the base independently of each other, somewhat 
recurved; cross veins numerous, forming a reticulation over much 
of the wing; intercalary sectors in radial and medial area. 
Relationships: The family Calvertiellidae clearly has a very 
specialized venation and seems far removed from most of the families 
of the order. This is shown by the short Sc, proximity of CuA and 
MP, the wide separation of CuA and CuP, the curved CuP, 
recurved anal veins (fore wing) and presence of intercalary sectors. 
Its closest relatives in the order seem to be the Eugereonidae 
(including Dictyoptilidae ; see Carpenter, 1964, p. 104), which differ 
from Calvertiellidae in having Sc long and terminating on C, CuA 
more remote from the posterior branch of MP, CuP branched and 
less arched, area between CuA and CuP narrower, anal veins less 
arched, absence of intercalary sectors and of the cuticular thickening. 
The hind wing of Calvertiellidae is broader than that in the 
Eugereonidae and more triangular in shape. 
It is interesting to find in the wings of the Calvertiellidae a short, 
simple or branched veinlet, which has now been noted in several 
families of Palaeodictyoptera (e.g. Breyeriidae, Kukalova, 1958; 
Spilapteridae, Carpenter, 1964). In Calvertiella it is a simple 
convex vein, but in the hind wing of Moravia n.g., it is better 
developed, with a series of short veinlets. Although this vein may 
actually be a basal branch of the costa, since the area set off cannot 
reasonably be considered homologous with the precostal area of the 
Orthoptera, I have termed it the postcostal area (Kukalova, 1963). 
The vein seems more likely to be homologous with the costal brace 
of Ephemeroptera, resembling especially the Permian types. 
Geological occurrence of family: Lower Permian of Kansas 
(USA) and Moravia (Czechoslovakia). 
Genera included: Calvertiella Tillyard, 1925; Moraviptera 
Kukalova, 1955; Moravia , new genus (herein). 
Genus Calvertiella Tillyard 
Calvertiella Tillyard, 1925, Amer. Journ. Sci, 10:43; Tillyard and Fraser, 
1938, Austral. Zool., 9(2) :141-142 ; Fraser, 1957, Royal Zool. Soc. 
N SW :24. 
