196 
Psyche 
[December 
hind wings are much less specialized than those of the Protelytridae. 
There seems to be little question on the basis of the evidence which 
Tillyard has given (1931) that the remigium of the protelytrid 
hind wing was capable of transverse folding at the level of the 
fulcrum; the hind wing is distinctly longer than the fore wing and 
the presence of the “hinges” on the anal veins, in line with the 
fulcrum, is indeed strong evidence. On the other hand, it is doubtful 
that the hind wings of the Blattelytridae were capable of transverse 
folding; indeed, the hind wings, as they are known to us in this 
family, are not as long as the front wings and could, therefore, have 
been completely covered by the elytra in the resting position without 
the transverse folding. In the case of the family Apachelytridae, the 
hind wing is slightly longer than the elytra but there are no signs 
of “hinges” on the veins and the fulcrum is not developed. The 
condition of the wings and wing venation strongly indicates that 
these three families are not part of a single evolutionary line within 
the order but represent three radiating lines. It is especially interest- 
ing to note in the Apachelytridae CuP and iA are not close together 
or parallel, as they are in the other two families. 
The body structure of the Protelytroptera is not sufficiently known 
to permit any general conclusions about lines of evolution within the 
order. The head apparently was relatively small with prominent 
eyes. The antennae prominent, relatively thick basally; known only 
in the Protelytridae and the Blattelytridae, they are about half the 
length of the fore wing. The prothorax apparently included a small 
pronotal disc. The prothoracic legs were apparently short, with five 
tarsal segments; the hind legs were much longer apparently with 
well developed femora and five tarsal segments, also. The abdomen 
terminated in a pair of short but distinctly segmented cerci ; the 
females (Apachelytridae) apparently possessed a short ovipositor. 
References 
Carpenter, F. M. 
1933. The Lower Permian Insects of Kansas. Part 6. Proc. Amer. 
Acad. Arts Sci. 68:4-11-503. 
1939. The Lower Permian Insects of Kansas. Part 8. Proc. Amer. 
Acad. Arts Sci. 73 :29-70. 
Martynov, A. V. 
1932. Permian Fossil Insects from the Arkhangelsk District. Part II. 
Trav. Inst. Paleozool. Acad. Sci. URSS. 2:63-96. 
Rohdendorf, B. B. 
1939. A new Protelytropteron from the Permian of the Urals. C. R. 
Acad. Sci. URSS. 23:506-508. 
