88 
Psyche 
[March 
Figures 1-3. Metaphase plates of dacetine species: (1) Orectognathus 
clarki, Ferntree Gully sample. Diploid cell from air dried preparation 
of pharate pupal cerebral ganglion (2) Epopostruma sp. Haploid cell 
from squash preparation of pupal testis. (3) Colobostruma alinodis. Dip- 
loid cell from air dried preparation of pharate pupal cerebral ganglion. 
Line in figure 3 represents 10 microns. 
in which all the chromosomes are metacentric, with a more or less 
continuous gradation in size. A lack of consistent differences pre- 
cluded grouping the chromosomes. 
Colobostruma alinodis (figure 3) 
n = 11, 2n = 22 
Material from Ferntree Gully State Park, Victoria, shows a 
karyotype with eleven metacentric chromosomes. As in the case of 
the Epopostruma karyotype above, there is a range in size of the 
chromosomes, but without consistent discontinuities enabling an ar- 
rangement of the chromosomes into groups. 
DISCUSSION 
In Brown and Wilson’s (1959) phylogenetic scheme for the Dace- 
tini, Colobostruma and Epopostruma are placed very close together, 
with Orectognathus some distance away. The cytogenetic results 
tend to support this placement although, even between the Colobos- 
truma and Epopostruma karyotypes, at least three changes must have 
occurred (one centric fusion or dissociation, and two pericentric in- 
versions). The difference between these karyotypes and that of 
Orectognathus clarki is substantial, and speculation about the changes 
involved would be idle. 
