1970] 
MacLeod — Baltic Amber Neuroptera 
159 
Pronymphes Kruger 
Pronymphes Kruger, 1923, Stett. Ent. Zeit. 84: 75-80. 
Type species (by original designation) : Nymphes mengeanus 
Hagen. Bachofen-Echt, 1949, Der Bernstein und seine Einschluse, 
P. 136. 
Description. Wings narrowly elongate. Fore wing: costal space 
narrow, not abruptly widened at base, costal crossveins unforked ; 
Rs+MA originating near base of wing, fused for a rather long 
distance before separation of MA; MP deeply forked at a level 
which is distinctly proximal to separation of MA from Rs, the two 
branches of MP running nearly parallel, the posterior branch sepa- 
rate from CuA ; basal branches of Cu running close together and 
nearly parallel, CuA unforked in portion preserved, CuP with 
pectinate branches to hind margin of wing 5 lacking crossveins 
between branches basally, perhaps with a series of irregular cross- 
veins distally. 
Hind wing: costal space and crossveins as in fore wing; origin 
of Rs + MA much further distal than in fore wing; MP deeply 
forked to base of wing, MP 2 with wide fork slightly distal to origin 
of Rs + MA ; Cu very short, with only a few pectinate branches to 
hind margin of wing, unconnected by crossveins. 
Discussion. The elongate, narrow form of the wings of Pro- 
nymphes is quite unlike the short, broad wings with abrupt basal 
dilations of the costal space to be found in the living genera 
Myiodactylus and N ymphidrion and I believe that these genera bear 
no close phylogenetic relationship to the fossil genus. There are 
such additional differences in venational details as the unforked 
MP in the fore wing of Myiodactylus to reinforce this conclusion. 
The wings of the species of Osmylops, though somewhat narrower 
in proportion to their width, are still distinctly broader than those 
of Pronymphes and the costal space of Osmylops also has a pro- 
nounced dilation near the base. 
In terms of their overall similarity in wing shape and venation, 
Pronymphes is obviously close to the living genera Nyjnphes , Aus- 
tronymphes , and, particularly, Nesydrion. Unfortunately, attempts 
to infer the probable details of the interesting phylogenetic inter- 
relationship of these four genera, from which I believe their phenetic 
similarities derive, will be somewhat frustrated until specimens show- 
ing the complete wings of Pronymphes are discovered. It is especially 
critical to learn the detailed structure of CuA in the fore wing and, 
particularly, whether this vein was forked or not. Certain similarities 
