THE MALE GENITALIA OF BLATTARIA. 
III. BLABERIDAE: ZETOBORINAE. 
By Louis M. Roth 
Pioneering Research Laboratory 
U. S. Army Natick Laboratories 
Natick, Massachusetts 01760 
In this paper I shall illustrate and characterize the male genitalia 
of 9 genera of blaberids which belong in the subfamily Zetoborinae. 
These genera are Capucina Saussure, Lanxoblatta Hebard, Phortioeca 
Saussure, Phortioecoides Rehn, Schistope/tis Rehn, Schizopilia Bur- 
meister, Tribonium Saussure, Zetobora Burmeister, and Zetoborella 
Hebard, Princis (i960) includes Tribonoidea Shelford in this sub- 
family but I have been unable to obtain a male of the one species in 
this genus. 
The male genitalia of most species of Blaberidae are characterized 
by the presence of three phallomeres: a median sclerite, a sclerite on 
the left, and a retractable hook on the right side. This led McKittrick 
(1964, p. 72) to conclude that “The male genitalia are extraordi- 
narily uniform throughout the Blaberidae.” She also stated (1964, 
p. 35) that “The male genitalia differ slightly in hook shape and 
the configuration of the posterior tip of the median sclerite within 
the Blaberidae, but otherwise show little variation within that group,” 
and on p. no “The great uniformity of blaberid male genitalia pre- 
cludes drawing many conclusions regarding subfamily relationships, 
except in the case of the Blaberinae and Panesthiinae.” “The great 
uniformity” of the male genitalia of Blaberidae is generally true only 
for the usual presence of three distinct phallomeres; in certain genera 
one or more of the phallomeres may be greatly reduced and even 
absent (e.g. R2 in Panchlora) . However, the shapes of the phallo- 
meres, especially the retractable hook (R2), prepuce, and L2d are 
often so distinctive that they may show subfamily, tribal, generic, 
species Group characteristics, and in some genera may be used for 
specific determination (Roth and Gurney, 1969; Roth, 1969, 1970). 
Materials and Methods 
The technique for preparing the genitalia has been described in pre- 
vious papers (Roth, 1969, 1970). The source of each of the specimens 
illustrated is given, using the following abbreviations: (ANSP) — 
Academy of Natural Sciences, Philadelphia; (BMNH) = British 
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