2 1 8 
Psyche 
[June 
Museum (Natural History), London; (CUZM) = Copenhagen 
University, Zoological Museum, Denmark; (MCZ) = Museum of 
Comparative Zoology, Harvard University; (N) = Natick culture, 
U. S. Army Natick Labs., Massachusetts; (USNM) = United 
States National Museum, Washington, D.C. 
Geographical collection data, and the names of specialists who iden- 
tified the specimens, if known, follow these abbreviations. The num- 
ber preceding the abbreviations refers to the number assigned the 
specimen and its corresponding genitalia (on a slide) which are de- 
posited in the museums indicated. 
Results and Discussion 
The basic features of the 3 phallomeres of the male genitalia of 
Zetoborinae are as follows: R2 (hooked right phallomere) is well 
developed and usually has a subapical incision (Fig. 32). The pre- 
putial membrane has a heavily sclerotized area (L2d) which extends 
dorsally or dorsolaterally to the right of L2vm (Fig. 28) ; L2d is 
flattened in the preparations and it should be remembered that it 
usually extends dorsally. Li is very well developed and various areas 
are darkly sclerotized (Fig. 30). 
Based on the degree of development of L2d and shape of L2vm, 
the Zetoborinae may be divided into 3 Tribes as follows: 
1. Zetoborini ( Zetobora and Phortioecoides , Figs. 1-5, 22-39). 
L2d is least developed in this tribe and is a thin sclerotization of the 
preputial membrane which extends dorsally (Figs. 22, 25, 27, 28, 
31, 34, 37). The subapical incision of R2 is clearly defined (Figs. 
23, 29, 32, 35, 38). Li is very similar in all the species in this 
tribe (Figs. 24, 26, 30, 33, 36, 39). 
Hebard (1921, p. 245) stated that “The differences shown by the 
forms lecognized by us as Tribonidium [ — Zetobora ] monasticum , 
transversum, and amplum f may indicate full specific distinction, geo- 
graphic racial differentiation or, indeed, mere individual variation in 
a single exceptionally plastic species.” The male genitalia of the 
above 3 species (Figs. 22-26, 28-36) are so similar that they are of 
little help in distinguishing the various species of Zetobora. 
Rehn (1937) placed Phortioecoides in the Panchlorinae. Princis 
(1961) followed Rehn and placed it in the Panchloridae. Princis’ 
Panchloridae is based principally on the shape of the subgenital plate 
in the male. However, the subgenital plate of the type male of 
Phortioecoides guarani Rehn (the only species of the genus) is not 
at all like that of Panchlora. Also the genitalia of Panchlora are 
