1970] 
Roth — * Blattaria 
233 
In Phortioeca peruana (Fig. 87) and P. phoraspoides (Fig. 98), 
the sclerotization of the preputial membrane, which forms the base 
of L2d, may be incomplete leaving a lacuna or window. However, 
it is clear that this character is variable (Figs. 90, 93, 96-98) and 
cannot be used for specific determination. In P. phoraspoides the 
rounded margin of the preputial membrane which surrounds the 
sclerotized base of L2d has more or less uniformly spaced indentations 
(Fig. 93) which are absent in other species of Phortioeca examined. 
Based on the shape of R2, Phortioeca nimbata , maximiliani , 
apolinari , and verrucosa can be grouped together. In these species 
the curved hook portion of R2 is relatively narrow and its subapical 
incision is poorly developed or absent (Figs. 73, 76, 79, 82, 85). 
Phortioeca peruana and P. phoraspoides both have stouter and more 
robust R2’s and the subapical incisions (Figs. 88, 94) are distinct. 
Rehn (1932) stated that the genera Zetobora , Lanxoblatta , Zeto- 
bofiella, and Schizopilia . . are clearly derivatives of a single 
phylum,” whereas Sckistopeltis and Tribonium . . typifies a dis- 
tinct and clearly marked phylogenetic series.” In general the male 
genitalia tend to support Rehn’s hypothesis. However, I have sep- 
arated Zetobora from the other 3 genera of his grouping because 
of the relatively poorly developed L2d. 
3. Triboniini ( Tribonium and Schistopeltis, Figs. 18-21, 99-113). 
In this tribe, L2vm has a well developed posterior arm or extension 
(Fig. 105) which extends dorsally (laterally in the photographs 
because of flattening) and L2d extends upward and curves towards 
this outgrowth (Figs. 99, 102, 105, 108, 111). The subapical in- 
cision of R2 is clearly defined and occurs at about the middle of the 
hook (Figs. 100, 103, 106, 109, 1 12). 
Summary 
Based on the shape of L2vm and L2d of the male genitalia, 9 
genera of Zetoborinae are divided into 3 tribes as follows: 
1. Zetoborini (Zetobora and Phortioecoides ) . 
2. Phortioecini ( Capucina , Lanxoblatta , Phortioeca , Schizopilia , 
and Zetoborella ) . 
3. Triboniini (Tribonium and Schistopeltis ) . 
