1970] 
Roth — Blattaria 
309 
to the subfamily Blaberinae. The genera Hyporhicnoda , Phoetalia , 
and Oxycercus, which were placed in subfamilies other than the 
Blaberinae by McKittrick and Princis, have male genitalia char- 
acteristics of members of Blaberinae and I include them in this 
subfamily. 
Materials and Methods 
The technique for preparing the male genitalia is given in Roth 
(1969). Unless otherwise indicated in the explanation of figures, 
the 2 phallomeres L2d (Fig. 41) and Li (Fig. 43) were mounted 
dorsal side up and the genital hook R2 (Fig. 42) is oriented ventral 
side uppermost. The source of each of the specimens illustrated is 
given using the following abbreviations: (AMNH) = American 
Museum of Natural History, New York; (ANSP) — Academy of 
Natural Sciences, Philadelphia; (BMNH) = British Museum 
(Natural History), London; (CUZM) — Copenhagen University, 
Zoological Museum, Denmark; (L) = Zoological Institute, Lund, 
Sweden; (LEM) = Lyman Entomological Museum, Macdonald 
College, Quebec, Canada; (N) = U. S. Army Natick Labs., Natick, 
Mass.; (USNM) = United States National Museum, Washington, 
D. C. Geographical collection data and the names of specialists who 
identified the specimens, if known, follow these abbreviations. The 
number preceding the abbreviations refers to the number assigned the 
specimen and its corresponding genitalia (on a slide) which are 
deposited in their respective museums. 
Results and Discussion 
The male phallomeres which have proved useful in studies of 
Blaberidae are a retractable hook on the right side (R2) (Fig. 42) ; 
a median sclerite (L2vm) which may or may not be solidly attached 
to a sclerite L2d (L2 dorsal) ; and lying below or partly surround- 
ing L2d is a membrane, the prepuce, which almost always bears 
spines of various sizes and shapes (Fig. 41). The phallomere (Li) 
on the left side, has a heavily sclerotized cleft (Fig. 43) and a 
well defined membranous lobe which overlaps the cleft area; because 
of its transparency this membrane is difficult to see in most of the 
photographs. 
Rehn and Hebard (1927, pp. 257-258), in discussing the Blabe- 
rinae, concluded that there are . . at least three lines of develop- 
ment or phyla, which are as follows: 
I. Archimandrite 1, Blaberus, Eublaberus 
