176 Psyche [June 
a dragline in a preferential direction if directional cues were de- 
pendent on the mechanical properties of the line. 
Summary 
The dragline-following behavior of male wolf spiders was ob- 
served in response to male, female and imitation draglines. Male 
L. rabida followed female draglines suspended above the sub- 
stratum but not those laid directly on the substratum. These males 
followed male silk and imitation lines if laid contiguously with 
female draglines, suggesting that they depend on tactile cues dur- 
ing dragline-following. On the other hand, male L. punctulata 
followed both aerial lines and ground lines laid by females but did 
not follow imitation or male lines. This suggests that male L . 
punctulata are more sensitive to tactile and/or chemical cues than 
are male L. rabida. Microhabitat preferences of the two species 
may explain the above differences. Data also suggest that males 
of both species are able to extract directional information from 
the dragline, perhaps by tensional cues. 
Acknowledgments 
The author wishes to express his gratitude to Dr. J. S. Rovner 
for his support, valuable discussions and assistance in the prepara- 
tion of the manuscript. Thanks are also due Dr. W. D. Hummon, 
Dr. C. P. Spirito and Dr. G. E. Svendsen for their helpful criticism 
of this work. Anne Tietjen assisted in the preparation of the man- 
uscript. 
Literature Cited 
Bristowe, W. S. and G. H. Locket 
1926. The courtship of British lycosid spiders and its probable significance. 
Proc. Zool. Soc. 22: 317 347. 
Conover, W. J. 
1971. Practical Nonparametric Statistics. New York: John Wiley and Sons, 
Inc. 492 pp. 
Dijkstra, H. 
1970. Comparative research of the courtship behaviour on the genus Pardosa 
(Arach. Araneae) III. Agnostic behaviour in Pardosa amentata. Bull. 
Mus. Nat. Hist. Natur. 41 : 91-97. 
Dondale, C. D. and B. M. Hegdekar 
1973. The contact sex pheromone of Pardosa lapidicina. Emerton (Araneida; 
Lycosidae). Canad. J. Zool. 51: 400-401. 
