1977] 
Peck — Leiodinae and Catopinae 
247 
Trelawny Parish. 5 mi N Alberttown, January, 1973, litter Ber- 
lese 250, LW females 3, LW males 8. Discovery Bay Marine Lab., 
10 feet, September, 1974, dung trap LW males 1. Drip Cave, 
August, 1974, SW females 6, SW males 6. Windsor Great Cave, 
March, 1973. R. Norton leg., SW females 8, SW males 10; August, 
1974, SW females 54, SW males 72. Windsor, 500 feet, August, 
1974, litter Berlese 290, SW females 3, SW females 1, SW males 2, 
SW males 3. 
These data show the species to be spread across the island in 
humid forests, from sea level to 7400 feet elevation. 
Discussion. For purposes of analysis of wing dimorphism, the 
above data has been combined with that of this species in my 1972 
paper. The data show that there is no particular relationship be- 
tween wing condition and sex. The sex ratio is statistically dis- 
tributed around an equal ratio of males to females, and the short 
wing form is not significantly more prevalent in either sex. 
However, there is a relationship between wing condition and 
habitat. In all 15 caves known to be inhabited by the beetle (rang- 
ing from 200 feet to 1750 feet in elevation), the populations are 
almost exclusively short winged. The exceptions are two and 10 
long winged specimens from Peru and Mocho Caves respectively. 
Otherwise, all the other 516 cave-collected specimens are short 
winged. 
A second relationship exists between wing condition and forest 
habitat elevation. Generally, the short winged form is more prev- 
alent at higher elevations. Combined data were analyzed to try to 
determine the existence of a quantitative relationship between wing 
reduction and elevation, but the scatter of the data points, even 
after an arc sin transformation, was too great to allow the calcu- 
lation of a meaningful regression line. This is because many of the 
lower elevation samples are too small to significantly record wing 
condition in the population. Even large Berlese samples often 
yielded only one or two specimens. Another error source is that 
pitfall traps in the lowlands selectively sampled long winged indi- 
viduals because they are more able to arrive at the traps by flight. 
The increased occurrence of wing reduction at higher elevations 
has been noted and discussed by Darlington (1943, 1970) for tem- 
perate and tropical carabid beetles. The phenomenon has also 
been an important component in the unraveling of the Pleistocene 
and Recent distributional history of carabids in Scandinavia (Lind- 
