298 
Psyche 
[September-December 
also began to accidentally dislodge legs of opponents, they could 
have evolved as described above to the present state, again losing 
their original function in the process. A combination of sequences 
1 and 2 could also have occurred. 
Summary 
The horns and long front legs of male Golofa porteri function 
as effective weapons in intraspecific battles between males. 
Acknowledgements 
I am indebted to Sr. Jorge Garavito of Telecom, Colombia for 
providing me with a crucial batch of beetles, and I thank Drs. 
M. J. W. Eberhard and H. Howden for criticizing the manuscript. 
This work was supported financially by the Comite de Investiga- 
ciones, Universidad del Valle. 
Literature Cited 
Eberhard, W. G. 
In press. The function of horns in the dynastine Podischnus agenor and other 
beetles. In: M. Blum and A. Blum (ed.) Sexual Selection and Repro- 
ductive Competition in Insects. Academic Press, New York. 
Howden, H., and M. Campbell 
1974. Observations on some scarabaeoidea in the Colombian Sierra Nevada 
de Santa Marta. Coleop. Bull. 28(3): 109-114. 
Joseph, C. 
1928. El Chiasognathus Grandti Steph. Rev. Universitaria (U. Catolica) 
(Santiago, Chile). 13(1): 529-535. 
Wille, J. E. 
1943. Entomologia Agricola del Peru. Ministerio de Agricultura, Lima. 
