1974 d Shear — Milliped Genus Bolhnanella 135 
Berlese extraction which Benedict and Malcolm have carried out in 
Oregon. 
It is now possible to do a number of things about Bolhnanella , 
which in 1970 I had to treat as a nomen dubium. 1) The genus can 
be characterized as fully distinct and valid, separate from, but related 
to, Taiyutyla and Conotyla, and more distant from Austrotyla and 
Achemenides , 2) some clear ideas of the distribution of the genus 
can be gained, and 3) seven new species can be described. 
The distribution of the eight known species of Bollmanella ranges 
from southern coastal Oregon to just above the Columbia River in 
Mason Co., Washington, with one species reported here from the 
Wallowa Mountains in extreme northeastern Oregon. The scattered 
nature of the localities suggests that species of Bollmanella , except 
for B. oregona, have small ranges, and we might expect many more 
to be discovered. In particular, the isolated mountain ranges of 
eastern Oregon need exploration. 
Ecologically, Bollmanella species have been collected exclusively 
from litter and duff derived from deciduous trees. Conifer duff and 
litter support species of Taiyutyla , though members of this latter 
genus are also found in deciduous duff, often syntopically with Boll- 
manella species. Although B. oregona ranges from sea level to around 
2000' elevation, most of the other species seem to have been collected 
from forests or parkland between 900' and 1500' elevation. This 
contrasts again with some species of Taiyutyla , which (unpublished 
data) are known from quite high elevations. Species of an unrelated 
chordeumid genus, Rhiscosomides (Shear, 1973), are found at low 
elevations and seem to favor coniferous forests. This ecological data 
also represents information from many collections which did not 
contain specimens of Bollmanella , thus clearly suggesting which habi- 
tats are most favorable. 
All type specimens of new species have been deposited in the 
Museum of Comparative Zoology, Cambridge, Mass. 
Superfamily Heterochordeumatoidea Pocock 
Family Conotylidae Cook 
Genus Bollmanella Chamberlin 
Bollmanella Chamberlin, 1941, Bull. Univ. Utah Biol. Ser. 6(5): 12. Type 
species, B. oregona Chamberlin, by monotypy and original designation. 
Diagnosis: With the characters of the family. Small (ca. 6-7 mm 
long) conotylid millipeds with typical form, distinct in gonopod 
morphology. Anterior gonopods simple, usually acuminate, but may 
