3 1 6 
Psyche 
[June 
families and the wing base morphology of the Palaeodictyoptera and 
Megasecoptera will be discussed elsewhere. 
The new material includes six new genera and ten new species 
of Elmoidae from the Lower Permian beds near Obora in Moravia, 
Czechoslovakia. The base of the wing is well preserved in one 
specimen only, Permodiapha carpenteri n.sp. (No. 8/1974), and 
incompletely preserved in seven other specimens. 
Since no definitive studies of the wing bases of the extant Paleop- 
tera [Odonata and Ephemeroptera] have been made, some difference 
of opinion exists about the interpretation of the basal sclerites. The 
interpretation of the diaphanopteran structures used in this paper 
should be considered only as tentative. The terminology employed is 
derived from the account of the wing base of Odonata by Tannert 
(1958), since the fore wing base in the Ephemeroptera has never 
been described in functional terms. For further comparison, the 
wing base structures of the related family Martynoviidae (un- 
described material) and the related order Palaeodictyoptera (Kuka- 
lova i960, 1969, 1970) have been used. 
Family Elmoidae 
Elmoidae Tillyard 1937: 82; Carpenter 1943: 56; Carpenter 1947: 27; 
Rohdendorf 1962: 69; Carpenter 1963: 249. 
Parelmoidae Rohdendorf 1962: 71. 
The family Elmoidae was originally assigned by Tillyard to the 
paleopterous order Megasecoptera because of the similarity in the 
venation pattern. This point of view was followed by Carpenter 
in his emendation of Elmoidae in 1943 and 1947, although he sub- 
sequently (1954) separated the Elmoidae, along with the Diapha- 
nopteridae and related families, into a distinct suborder (Parame- 
gasecoptera) , on the basis of the folded, resting position of the 
wings. Later, Rohdendorf (1962) and Carpenter (1963) independ- 
ently reached the conclusion that these families had to be referred 
to the separate order Diaphanopterodea, established by Handlirsch 
(1919). A detailed evaluation of the phylogenetic position of Elmoi- 
dae and Diaphanopteridae within the order, and of Diaphanopterodea 
within the Paleoptera has been given by Carpenter (1963). 
Until now, the family has been known by only three genera, all 
from the Lower Permian of Kansas and Oklahoma — Elmoa, 
Parelmoa and Pseudelmoa, represented by five species. The new 
fossils from Moravia contribute additional and varied morphological 
features, which broaden the family characteristics formerly used. In 
the Moravian material thickened cuticular spots are sometimes pres- 
