396 
Psyche 
[September-December 
the Cumberland Plateau of Kentucky and Alabama, the Highland 
Rim of Tennessee, and the Shawnee Hills of southern Illinois are 
probably not remnants of an ancient distribution but also date from 
the Wisconsinan glaciation when more wet and cool montane-type 
climatic conditions prevailed in the Interior Basins, favoring the 
distribution and dispersal of “montane” beetles. 
The comparatively large ranges and low levels of geographic vari- 
ability in C. ulkei and C. appalachianus are remarkable in the light 
of an assumed low dispersal potential resulting from the beetles’ 
winglessness, eyelessness, and soil habitation. Other ecologically 
similar beetles, for instance Anillinus and Arianops (Barr, 1969; 
1974) in the eastern United States have much smaller ranges and 
are more highly speciated. In contrast then, it must be that the 
vagility of some Catopocerus is higher than would seem likely at 
first, because of an incompletely understood ability to withstand 
various mechanisms of long distance dispersal, similar to that of some 
European soil Colydiid beetles (Peck, 1972). 
Acknowledgements 
I thank the following persons and their institutions for allowing 
examination of specimens: M. G. Emsley, ANSP; L. H. Herman, 
AMNH ; H. B. Leech, CAS; J. M. Campbell, CNCI; H. Dybas 
and R. Wenzel, FMNH ; M. W. Sanderson, INHS; P. J. Darling- 
ton and J. F. Lawrence, MCZ; J. M. Kingsolver, USNM; Fr. 
Jerome Rupprecht, SVAM ; G. E. Wallace, CM; and G. Byers, 
UK. Thomas C. Barr, University of Kentucky, and Henry F. 
Howden have provided specimens from their personal collections. 
Collections made in regions administered by the National Park Ser- 
vice were permitted by Ernest G. Whanger, Blue Ridge Parkway, 
and V. R. Bender, Great Smoky Mountains National Park. A debt 
of gratitude is due to many persons who assisted me in collecting, 
especially James H. Peck in 1965, Alan Fiske in 1967, and Jarmila 
Kukalova-Peck. Dr. and Mrs. Walter B. Jones of Huntsville, Ala- 
bama, are thanked for their hospitality in providing a base for field 
work conducted intermittently in and near Alabama over seven years. 
My wife, Jarmila, prepared the final illustrations. My specimens 
were collected during field work supported by National Science 
Foundation Grants GB 3167 and GB 7346 (Professor Reed C. 
Rollins, principal investigator, Biological Laboratories, Harvard Uni- 
versity) and by Canadian National Research Council operating 
grants. John Lawrence and Ernst Mayr kindly read an early ver- 
sion of the manuscript. 
