450 
Psyche 
[September-December 
northern Sierran (Coloma-Aukum) population than to the Mariposa 
population. 
Noteworthy geographic variation also occurs in thoracic groove 
shape, pedipalpal femur shape, and conductor form. Almost all 
coastal specimens have an elongate thoracic pit which narrows pos- 
teriorly (Fig. 45). The Coloma-Aukum specimens have roughly 
circular or slightly transverse pits. The Mariposa specimens have 
slightly elongate pits which narrow posteriorly and have a roughly 
triangular shape. Coastal specimens have rather strongly bowed 
pedipalpal femurs (Fig. 79), Mariposa pedipalpal femurs are less 
strongly bowed, and Coloma-Aukum ones are nearly straight (Fig. 
78). Most of the Sierran specimens have less well developed ICS 
keels and relatively wider conductor tips (Figs. 98-100) than do 
coastal specimens (Figs. 96-97). However, there is considerable 
variation among Sierran populations, with Coloma males having a 
rather well developed keel and narrow conductor tip (Fig. 98), and 
Mariposa males having smaller keels and proportionately wider con- 
ductor tips (Figs. 99-100). 
Females: There is a moderate amount of geographic variation 
among the coastal populations. Although some individual population 
samples (especially the Mt. Diablo and Soquel area samples) are 
quite different from another sample in a few ratios (Figs. 16-17), 
no population sample is distinct from the rest of the entire coastal 
sample in any character. The three northern Sierran (Coloma- 
Aukum) specimens are quite similar to the coastal populations in all 
characters with the exception of seminal receptacle form. Variation 
in the number of stalk bends in the northern Sierran receptacles 
(Figs. 1 27- 1 29) spans the gap between the more sinuous coastal 
stalks (Figs. 1 21-125) and the less sinuous Mariposa stalks (Figs. 
130-131). Northern Sierran females tend to have proportionately 
large receptacle bulbs (Figs. 128-129). The Mariposa sample 
differs rather strongly from the coastal samples in four characters 
(Figs. 13-15) : IFL/IVFL, IVFL/IVTL, CL/IFL, and CL/ITL. 
It likewise differs markedly from the northern Sierran sample in 
these plus a fifth, CL/IVTL (Fig. 16). 
In summary, male and female variation patterns indicate that there 
is little, if any, gene flow between coastal and Sierran populations 
across the unfavorable Central Valley, that there is little gene flow 
between the northern and more southern Sierran populations, and 
that the coastal population is genetically more similar to the northern 
Sierran populations than to the Mariposa population. I do not feel 
that the variation discontinuities between coastal populations and the 
