Psyche 
[Mar. 
tendency to limit foraging to more and more lengthy intervals, 
feeding the larvae less and less upon insect prey and more and 
more upon ingluvial food, until, at last, the habit of foraging 
disappeared altogether. 
The higher tribes among the Ponerinae, especially the Ecta- 
tommini, the Odontomachini, and the Ponerini are of peculiar 
interest in this connection. Here two of the three requisite 
physiological specializations, at least, seem to be moderately 
developed in certain species. Considerable discrepancy of stature 
between the perfect female and the first-brood workers is notable 
in a number of species of all these tribes. The power of regurgi- 
tation, though feebly developed, has been reported for the Bra- 
zilian Odontomachus affinis by Borgmeier (1920), for Ecta- 
tomma tuberculatum by Cook (1904— 5) and has been observed 
in Euponera gilva harnedi by Haskins (1931). It might be 
supposed, therefore, that among these higher Ponerinae, exam- 
ples could be found in which the females, unusually capable of 
fasting, tended to supply the first-brood larvae at least par- 
tially with ingluvial salivary or regurgitated substances and 
only in part with captured insect food, and to forage much less 
frequently than the fertile females of Myrmecia or Amblyopone. 
A very definite suggestion of this intermediate situation was 
obtained in the course of the present study with Odontomachus 
haematoda. A young female, taken immediately after the nuptial 
flight and dealation, isolated itself in the artificial nest in a 
closed cell in the typical manner reported by a number of stu- 
dents of this genus. In this case, however, all opportunities to 
obtain nourishment outside the nest were withdrawn, so that the 
young queen was thrown entirely on its own resources. Under 
these conditions, this Odontomachus female produced numerous 
eggs, hatched them, and evidently fed the resulting larvae with 
ingluvial food (although the process was difficult to observe) 
since they developed rapidly. They remained healthy for some 
time but, when somewhat less than one-half mature, declined 
slowly and eventually perished. Shortly after this, the female 
perished also. It had been captured on December 9, 1947, and 
died on March 3, 1948. Thus, although it had been unable to 
bring its young larvae to mlaturity or to establish a permanent 
formicary without outside nourishment, it had raised a good- 
sized brood through almost half the period of growth on nour- 
