1969] 
Illavac — Scarites 
9 
width 1.50. Total length (including peduncle) 21.8. Specimen ex- 
amined: holotype. 
Comments : known from the appendageless fragments of two speci- 
mens. I regard it as of uncertain subgeneric position for three reasons. 
The structures containing the major diagnostic characters of Antil- 
Uscaris, antennae and metepimeron, are absent in the available ma- 
terial. A major similarity between Antilliscaris and darlingtoni is 
the reduction of the flight mechanism. Flightlessness and, in extenso , 
large-scale reorganization of the flight mechanism occur with high 
frequency among montane carabids (Darlington 1943), which argues 
against uniting darlingtoni with Antilliscaris in the absence of other 
evidence. Furthermore, there is a, gap between Antilliscaris and dar- 
lingtoni in the dorsal surface of the mandibles (Figs. 5, 6, 7, 10, 11). 
Discussion 
Several character states in Antilliscaris range from atypical to 
unique for a Scarites s. 1 . Many of these differences are found in 
the locomotory system or are peripheral to it. This raises questions 
about associated behavioral and ecological differences. In an attempt 
to list some of the possible answers to these questions, particular 
configurations of Antilliscaris are contrasted with S. (s. s.J sub- 
striatus Haldemann and subterraneus Fabricius. This is not done 
because I feel that these 2 species are closely related to Antilliscaris, 
but because both fly and burrow (Hlavac, 1967) and can be used as 
a base line in interpreting the morphology of Antilliscaris . 
antennae: The antennae of Antilliscaris are unusual in three 
respects: in length, in the setation of the 4th segment and in the 
relative narrowness of the lateral glabrous areas. An increase in the 
size of the antennae can be documented in terms of elongation of 
individual segments, as expressed by length/width^ and in terms of 
increase relative to body size. The ranges of length/width of seg- 
ments 6-10 is 0.9-1. 1 in danforthi , 1. 35-1. 55 in substriatus , 1 .3-1.6 
in mutchleri, and 1.8-2. 3 in megacephalus. The antennae reach the 
procoxae in substriatus, the posterior margin of the prothorax in dan- 
forthi , and middle of the peduncle in mutchleri and the first abdominal 
sternite in megacephalus. As compared to substriatus , the antennae 
of danforthi are longer and the individual apical segments are much 
wider. Assuming that there are no major differences between the rel- 
ative widths of the first segment in the above species, then a measure 
of width of the antennal apex can be expressed as width of an apical 
segment/width of segment 1. Using width of the 8th segment in this 
ratio, the figures are 0.72 for substriatus and megacephalus, 0.93 for 
mutchleri and 1.1 for danforthi. 
