1969] 
1 Hlavac — Scarites 
1 1 
The elongate and/or broad apical segments and the state of the 
4th segment in Antilliscaris represent an increase in surface area of 
antennal apex over substriatus. Due to differences in distribution of 
surface types, there is a large increase in setose areas, which can be 
interpreted as an intensification of the functions served by this region. 
eyes: The eyes of Antilliscaris are small, 30-50% the height of 
the mandible, vs. 100% in substriatus , and are nearly circular except 
in danforthi where the posterior border is nearly straight. Smith 
(1964, p. 1 16) stated that the eyes of flightless carabids are not 
reduced, except for cave inhabiting species; this is clearly not the 
case for montane Scarites or for the montane carabid fauna of New 
Guinea (P. J. Darlington Jr., personal communication). Eye re- 
duction in Scarites seems to have come about either from a reduction 
in diameter (Antilliscaris) or from diameter reduction and shortening 
the anterior posterior axis, giving the eyes of S. (Typhloscaris) and 
S. (Taeniolobus) cubanae Banninger a peculiar slit-like appearance. 
mandibles: The mandibles of scaritine carabids are structurally 
complex and can be divided into three functional regions, an apical 
and 2 medial carnassal areas, and a basal molar part. Antilliscaris 
and Scarites s. s. represent extremes in the relative sizes of the latter 
2 of these areas (Figs. 14, 15). In an unworn specimen of sub- 
striatus (Fig. 5) the apical carnassal is 34%, the median carnassals 
26% and the molar 40% of the total length (perpendicular distance 
between apex and base) ; while in Antilliscaris (Figs. 6, 10, 11) the 
apical carnassal ranges from 24-34% the median carnassals from 
14 15 
Fig. 14. — Mandibles, S. (Antilliscaris). Fig. 15 — Mandibles, £. (s. s.). 
Medial carnassal areas shaded. 
