1969] 
•. Hlavac — Scarites 
17 
+ winged. A second montane species, O. reichoides Dari., occurs on 
Hispaniola. 
The sophisticated taxonomy necessary to separate monophyletic 
from polyphyletic origins of —wings in closely related lower cate- 
gories does not now exist for Scarites. Until a more refined sys- 
tematics becomes available, I think it heuristic and economical to view 
the biogeography of Antilliscaris in light of the model presented above. 
preliminary evolutionary diagnosis: From an assumed low- 
land, winged and burrowing ancestor, Antilliscaris has entered the 
montane rain forests of Puerto Rico, become flightless, undergone an 
ecological shift from burrowing to ground crawling, and speciated. 
References Cited 
Banninger, M. 
1935. Uber alte und neue formen der Scaritina. Ent. Blatt. 1935: 148- 
160. 
1937-39. Monographie der subtribus Scaritina (Col. Carab.). Pt. I — 
Deutsche Ent. Zeitschr. 1937: 6-160. Pt. II — Deutsche Ent. 
Zeitschr. 1938: 41-181. Pt. Ill — Deutsche Ent. Zeitschr. 1939: 
126-161. 
1949. Uber Carabinae. Mittellungen der Miinchner. Ent. Gesellschaft. 
Darlington, P. J., Jr. 
1939. West Indian Carabidae V. New forms from the Dominican Re- 
public and Puerto Rico. Mem. Soc. Cubana Hist. Nat. 13: 79-101. 
1941. West Indian Carabidae VI. The Jamaican species and their 
wings. Psyche 48: 10-15. 
1943. Carabidae of mountains and islands: data on the evolution of 
isolated faunas, and on atrophy of wings. Ecol. Monog. 13: 
37-61. 
Hlavac, T. F. 
1967. Observations on behavior in Scarites. Col. Bull. 21: 18-22. 
Larsen, O. 
1966. On the morphology of a function of the locomotor organs of the 
Gyrinidae and other Coleoptera. Opus. Ent. Supp. 30: 1-241. 
MacArthur, R. H., and E. O. Wilson 
1967. The theory of island biogeography. Princeton U. Press, 203 pp. 
Mutchler, A. M. 
1934. New species of Carabidae from Puerto Rico. Amer. Mus. Nov. 
No. 686, 5 pp. 
Smith, D. M. 
1964. The structure and development of flightless Coleoptera: a light 
and electron microscopic study of wings, thoracic exoskeleton and 
rudimentary flight musculature. J. Morph. 114: 107-184. 
