1969] 
Shear — Cave Millipeds 
135 
Jersey Co., Illinois 2 (Hoffman, 1962). Subsequent collections have 
shown it to be not uncommon in many of the caves of western Illinois, 
particularly in Adams, Pike, and Monroe Counties (S. Peck, pers. 
comm.). Its closest relatives occur in the Pacific Northwest, thus 
presenting another instance of the close faunistic correspondence be- 
tween the Ozark-Appalachian system and the Pacific coast mountain 
ranges. The case of E. remington i is remarkably similar to that of 
the antrodiaetid spider genus Atypoides, with a single species in Mis- 
souri and Illinois and two in California and Oregon (Coyle, 1968). 
Missouri caves should be thoroughly searched for nearctodesmids. 
ORDER SPIROSTREPTIDA 
Family Cambalidae 
The criteria for the establishment of genera, in this family are as 
yet uncertain. Loomis (1938) recognized several genera, most of 
them from the western United States, without resorting to details of 
the gonopod structure. More recently, Hoffman (1956) established 
the genus Troglocambala , for specimens from Turk’s Cave, Conecuh 
Co., Alabama, separated from Cambala by the three-jointed telopodite 
of the anterior gonopods. Causey (1964) implied without explana- 
tion that this genus was a synonym of Cambala , but established at 
the same time the genus Mexicambala , also a troglobite, based on 
specimens from Cueva de la Parra, San Luis Potosi, on differences 
of a similar quality. Until a thorough study reveals the true relation- 
ships of species in this family, it is my opinion that both these genera 
should be regarded as synonyms of Cambala. 
Genus Cambala Gray 
Fig. 1 
In the eastern United States, both Cambala annulata (Say), a 
large (up to 45 mm long), black species, and C. minor Bollman (Fig. 
1), a small (up to 20 mm long) brown species are troglophilic. The 
status of C. minor and a few other names based on similar forms is 
uncertain. Specimens from widely scattered caves in Alabama, Ten- 
nessee, Kentucky and West Virginia which I have recently exam- 
ined show small but constant differences in the gonopods, ocelli 
number, and pigmentation. Careful study will probably reveal that 
more than one species is involved. 
Cambala loomisi (Hoffman) has already been mentioned; it is the 
type species of Troglocambala. In Texas, the caves of the Edwards 
2 Now known as Grafton Cave. 
