1969] 
Kukalova — Palaeodictyoptera 
169 
mian, Czechoslovakia) ; Doropteron G. Zalessky, 1946 (Lower Per- 
mian, Ural, USSR) ; P ermiakovia Martynov, 1940 (Lower Permian, 
Ural, USSR) ; Abaptilon G. Zalessky, 1946 (Stephanian, Kuznetsk). 
The genus Hornaloneura occurs in the Westphalian of Illinois as 
well as in the Commentry shales (Carpenter, 1964, p. 117ft). 
Genus Hornaloneura Brongniart 
Hornaloneura Brongniart, 1885: 66; Brongniart, 1893 : 316 ; Handlirsch, 
1906: 107; Lameere, 1917: 148; Carpenter, 1964: 119. 
Homaloneurina Handlirsch, 1906: Lameere, 1917: 148; Handlirsch, 1919: 
20 . 
Homaloneuritcs Handlirsch, 1906: 107; Lameere, 1917: 147; Handlirsch, 
1919: 20. 
Type species: Hornaloneura elegans Brongniart, i885(OD). 
Although Brongniart originally erected Hornaloneura for a single 
species (elegans), he added five more species in 1893: bonnieri , 
punctata , joannae , ornata , and bucklandi. In 1906, Handlirsch 
formed two new genera, their type species being two of those de- 
scribed by Brongniart: he established Homaloneurina for bonnieri 
and Homaloneurites for joannae. Both of these genera, were rejected 
as unnecessary by Lameere (1917), a view which is undoubtedly 
correct and which is followed here. An additional species, lehrnani , 
is described below; this is based upon Brongniart’s specimen 17-15, 
which Brongniart erroneously considered to be the reverse half of 
the type specimen of ornata. 
The basic venational pattern within Hornaloneura is uniform, and 
many morphological features seem to have appeared independently 
among the species. It is usual for the species of Hornaloneura to have 
cuticular thickenings, supporting cross veins, and color bands in the 
basal third of both pairs of wings. These structures presumably 
strengthen transversely the thin wing membrane. Among palaeop- 
terous insects, they are most pronounced in the Odonata, but they 
occur in other orders as well. In the Palaeodictyoptera, they are most 
spectacularly represented by the Calvertiellidae (Kukalova, 1964). 
The strengthening structures in the wings of Hornaloneura were 
noted originally by Brongniart ( 1893, p. 318) , but neither Handlirsch 
nor Lameere made mention of them. In Hornaloneura elegans , bon- 
nieri } parva , and dabasinskasi , the cuticular thickening is a conspic- 
uous V-shaped ridge with its apex on Ai. In ornata , joannae ^ and 
lehmani , the cuticular ridge is directly on Ai, strengthening it for a 
short distance where it abruptly bends toward the posterior margin. 
Present in all species is a long, oblique, strong cross vein, running 
