70 
Psyche 
[June 
from M to R, usually at the point of origin of Rs. In elegans and 
bonnieri , there is additional strengthening by means of a dark colored 
band running obliquely from Ai to R. 
The body structures of Homaloneura were figured very roughly 
by Brongniart, and, with the exception of the prothoracic lobes and 
legs, they were not mentioned by Lameere. Actually, most of these 
structures are visible only under glycerin or glycerin-alcohol, but 
with the use of this clearing solution, the specimens of Homaloneura 
have contributed a great deal to our knowledge of the morphology of 
the Palaeodictyoptera. Most important is the presence of a large, 
bulging clypeus, with transverse ridges, best preserved in PI. lehmani , 
which I first noted when I examined this specimen under glycerin in 
1966. The structure of the clypeus is surprisingly like that of some 
Homoptera, such as the Cicadidae, and it undoubtedly indicates that 
in the Palaeodictyoptera a cibarium was present, much as in the true 
bugs. This was presumably developed in connection with the sucking 
beak. In the specimen of lehmani , the sheath of the beak is bent to 
one side, but the stylets are close together, not separated. 
Another definite morphological feature of the species of IPoma- 
loneura is the prothoracic lobes. These were presumably homologous 
with wings, but in any event, they show longitudinal veins as well 
as cross veins, and they are attached to the prothorax along a short, 
cuticular ridge, corresponding to the articular region of the meso- 
and metathoracic wings. The prothoracic lobes are cordate and more 
or less sclerotized, the veins showing no convexities or concavities. 
The lobes were somewhat higher on the thorax than the meso- and 
metathoracic wings, and there was apparently some space between 
them and the front wings in many species. The size of the lobes 
varies in the species of the genus. In those species in which the lobes 
were large, they apparently overlapped to some extent the fore wings. 
Since there is no indication of articular plates at the base of the lobes, 
active movement of the lobes in these Palaeodictyoptera is elim- 
inated. There is a possibility, however, that these lobes might have 
functioned as vanes in directing the flowing or movement of air in 
relation to the moving of the front wings. 
The abdomen in Plomaloneura was relatively narrow. Brongniart 
was of the opinion that only nine abdominal segments existed in the 
genus Homaloneura (1893, p. 316). This conclusion was apparently 
based on a specimen of ornata, the only Commentry specimen with 
the abdomen completely preserved, in which segments 1-3 are shorter 
than the following ones, and in which the posterior margin of the 
