1969] 
Kukalova — P alae o diet y opt era 
177 
Cross veins few, almost regular. Veins and cross veins lightly colored. 
Hind wing: length 28, width about 11 mm. 
The venation indicates that punctata is related to the elegans- 
bonnieri group more closely than to the j oannae-ornata-lehmani group, 
since the fore wing lacks the very oblique position of branches of 
CuA, and the main veins are more richly branched than in the last 
group of species. 
Homaloneura joannae Brongniart 
Figure 6 
Homaloneura joannae Brongniart, 1893: 320, pi. 18, fig. 5; Lameere, 1917: 
147. 
Homaloneurites joannae Handlirsch, 1906: 107, pi. 12, fig. 1. 
This species was based by Brongniart upon specimen 18-5, which 
consists of the four wings, thorax, one prothoracic lobe, and vague 
outlines of head and clypeus. Handlirsch (1906) erected for this 
species a separate genus, Homaloneurites , on the basis of its veins 
being less branched ; but that genus was placed in synonymy of 
Homaloneura by Lameere (1917), who did not accept as valid the 
characteristics given by Handlirsch. This is a difficult problem, 
and it is discussed below. 
Handlirsch (1906) thought that Brongniart’s specimen 17-15 was 
joannae as well, this fossil being erroneously designated by Brong- 
niart as the reverse half of the type of ornata (17-16). Handlirsch 
was right that the specimen (17-15) was not the counterpart of the 
type specimen of ornata ; it is not joannae either, but represents an 
undescribed species, named below lekmani. 
In Brongniart’s account of the specimen of joannae there are 
several mistakes in venational interpretation and in the description 
of the color pattern; also, the hind wing and the body structures 
were not illustrated. The following account gives these additional 
details as well as a review of those noted by Brongniart. 
Wings equally long, membrane relatively strong and dark, with 
transverse light bands. Cuticular thickening extending on Ai for 
a short distance as the vein curves towards the posterior margin. 
Supporting cross vein running obliquely from the stem of M to the 
very base of Rs. Fore wing: length 22.5 mm, width 5.7 mm. Fore 
wing broadest at the beginning of the apical third, narrowing to- 
wards the base; apex rounded, about on the wing axis; anterior 
margin slightly concave; postcostal area triangular and small. Rs 
with seven simple branches; MA forked 1-2 times; MP with a long 
fork; CuA with 4-5 pectinate branches; CuP simple; cubital branches 
