1969] 
Kukalova — Palaeodictyoptera 
195 
gin somewhat undulated. Hind wing little known, not greatly 
broadened. 
Body structures: Head small, much narrower than the prothorax, 
eyes prominent. Prothoracic lobes cordate, with radiating veins and 
numerous cross veins; metathorax very broad, longer than the meso- 
thorax. 
This genus is apparently related to other genera of this family 
with numerous cross veins, such as Palaeoptilus , Spiloptilus , Epitethe , 
and Becquerelia. The absence of the distal part of Ri in grehanti 
prevents precise comparison with Palaeoptilus , and the absence of 
the cuticular thickening of the wing in Palaeoptilus , Epitethe and 
Spiloptilus prevents satisfactory comparison with those genera. How- 
ever, the genus Tectoptilus differs from all other spilapterid genera 
in having the Y-shaped cuticular thickening, which extends between 
CuP and Ai, and in having an unusually broad area of CuP, with 
many branches. This last character, as well as the fully developed 
MA and the presence of numerous cross veins resembles the structure 
of the Fouqueidae, although the other features known are spilapterid 
in nature. The genus Tectoptilus , in this way, seems to be somewhat 
intermediate between the Fouqueidae and the Spilapteridae. At 
the present state of our knowledge, it seems to be impossible to draw 
a strict line between these two families. 
Tectoptilus grehanti (Brongniart) 
Figures 16 and 17 
Becquerelia grehanti Brongniart, 1893: 359, pi. 20, fig. 4; Handlirsch 1906: 
106, pi. 11, fig. 25; Lameere, 1917: 152. 
This species, based upon specimen 20-4, shows two hind wings, 
the metathorax, and five segments of the abdomen ; in addition, 
another, previously undescribed specimen is contained in the collec- 
tion of the Museum, this showing the fore wings in part, the head 
(with eyes and clypeus), prothoracic lobes, and the thorax and six 
segments of the abdomen. On the basis of the similarity of venation 
and the nature of the cuticular thickening, I consider this conspecific 
with the type of grehanti ; this may be the specimen which Lameere 
(1917: 155) mentioned as an additional specimen of Fouquea need- 
hami Lameere. The following account is based upon both of these 
specimens. 
Wings: dark with light maculations; fore wing: length 58 mm, 
width 20 mm, broadest slightly before the midwing; narrowed in the 
basal part with distal part elongate; r-rs area not broadened, with 
