1969] 
Roth - — Genitalia of Blattaria 
225 
Blaberus has this character, although I have seen a few very minute 
hairs in some specimens of B. giganteus. 
In craniifer the tumorlike sclerotized outgrowth on the left side 
of L2 extends caudally for a short distance and usually merges 
gradually into the border of the prepuce where small spines begin 
and form a fringe around the membrane. The sclerotized extension 
of the lateral outgrowth varies somewhat in length but is generally 
distinct (Figs. 47-51, 53-57, 62-64, 66-74). Exceptions are seen in 
Fig. 52 and 65. In B. giganteus there is usually little or no sclerotized 
extension from the tumorlike outgrowth into the preputial membrane, 
so that the preputial spines begin more abruptly at the outgrowth 
(Figs. 28-35, 38-40). Exceptions to this are shown in Fig. 36 and 37. 
In both species the preputial spines may occur in more than a single 
row, and in some individuals there may be a reduction in the number 
of preputial spines (Figs. 37, 65, 70, 74). 
If Princis’ (1946) tegminal character is valid for B. craniifer then 
I have not seen any correctly determined specimens of B. trapezoideus. 
All the specimens determined by Hebard or Rehn as trapezoideus 
(Figs. 41, 58-61) have hairy tegmina and I therefore consider them 
to be light forms of B. craniifer. The genitalia of these “trapezoideus” 
(e.g., Figs. 47, 63, 64, 66) are indistinguishable from craniifer. Two 
specimens identified as B. trapezoideus , received from the University 
Zoological Museum, Copenhagen, Denmark were actually B. craniifer 
(Costa Rica) and B. parabolicus (Peru). According to Princis 
(1946) the preputial spines of trapezoideus are similar to giganteus 
but are smaller and more numerous, though always clearly separated 
from each other. Considering the variation in size, number, and 
spacing of preputial spines I doubt if this character can be used to 
distinguish trapezoideus from giganteus. Princis also (1958) states 
that the pronotum of trapezoideus is laterally truncated with approxi- 
mately parallel sides. Some individuals of our light-phased form of 
craniifer (in culture) (Fig. 42) also have the pronotum laterally 
truncate. It is possible that trapezoideus and craniifer are simply 
variants of the same species. The type localities of the 2 spp. of the 
(from specimen shown in Fig. 43). 49. (USNM), Vera Cruz, Mexico. 50. 
(USNM), Tuxtepec, Oaxaca, Mexico (labelled trapezoideus) . 51. (USNM), 
Mexico (from specimen shown in Fig. 44). 52. (MCZ), Chichen Itza, Yuca- 
tan, Mexico. 53. (USNM), El Salvador. 54. (USNM), San Salvador, El 
Salvador. 55. (USNM), Benque Viejo, British Honduras. 56. (MCZ), 
Colon, Panama. (Abbreviations for Fig. 52. L2d = dorsal left phallomere; 
L2vm = ventromedial left sclerite (fused to L2d) ; P — prepuce; T = 
tumorlike outgrowth.) 
