1969] 
Roth — Genitalia of Blattaria 
243 
by him with reservations) but he did not remove the genitalia from 
the male; it differs from Princis’ description in having a 4-pronged 
elevation on the left and more spines on the right (than he figured) 
terminating anteriorly in 3 spines fused at their bases (Fig. 204). 
Hebard stated that the species related to B. fusiformis Walker 
. . are poorly understood and the description of fusiformis is vague. 
If our specimens are correctly determined it is possible that anisitsi 
is a synonym, based on material showing decided depauperation.” 
Although Hebard included anisitsi in the Brasilianus Group, the 
specimen identified by Princis as anisitsi (Fig. 202) is clearly a mem- 
ber (by male genitalia, Fig. 203) of the Atropos Group (Princis 
1946). If both Princis and Hebard are correct in their determina- 
tions anisitsi and fusiformis are obviously not the same species. 
Princis (1946) compared the genitalia of boliviensis (Figs. 200, 
201) with anisitsi (Figs. 203, 204). According to him the penis in 
boliviensis is more massive. The spines on the right side start with a 
bluntly rounded spine, are larger, more numerous and not as widely 
separated as in anisitsi. On the left side there is a 3-pronged truncate 
elevation followed by 10 (according to his drawing) truncate or 
rounded spines set fairly close together. In the specimen shown in 
Figs. 200, 201 (not the one illustrated by Princis), the truncate ele- 
vations on the left are at least 6-pronged. No doubt an examination 
of additional specimens of these 2 species would show as much intra- 
specific variation as occurs in other species of the Atropos Group. 
The truncate elevations on the left side of both anisitsi and bolivien- 
sis arise close to L2vm, extend dorsally, and their genitalia closely 
resemble those of B. discoidalis . 
Undetermined species. — Several museum specimens were examined 
whose genitalia and phenotypic appearance did not fall into the known 
species. These were as follows: 
1-2. Blaberus spp. A (Figs. 205, 213) and B (Figs. 206, 21 1, 
212). — These 2 species from Peru, except for their much smaller 
size, resemble colosseus, particularly in their slender form and pale 
coloration. The preputial spines of both forms (cf. Figs. 21 1, 213) 
differ from each other. The anterior elevations of the left side are 
not much larger than those on the right, thus resembling the prepuce 
of colosseus. 
3. Blaberus sp. C (Fig. 207). — This specimen from Colombia 
was identified by Hebard as B. discoidalis. However, it is considerably 
smaller and more slender than is discoidalis , and phenotypically re- 
sembles the specimen identified by Princis as B. anisitsi (Fig. 202). 
Its genitalia (Fig. 210) are unique (and differ from anisitsi , Fig. 
