1969] 
Ponomarenko — Cretaceous Insects 
309 
between A4 and the sutural margin. It is distinguished by the com- 
plete reduction of CuP and by having all veins parallel to the elytral 
margin. 
Labradorocoleus, gen. nov. 
Type species: Labradorocoleus carpenteri, sp. nov. 
Epipleural space narrowing to apex. The rs2-area, is narrow 
before the middle of the elytron, with one row of cells; two rows 
present in the distal part of the elytron. The rest of the areas with 
two rows of cells. A3 more than a quarter the length of the elytron, 
the second row of cells in its area very short. Cells in the 24-area 
typically large. 
This genus includes a. single species from Cretaceous deposits of 
Canada. 
Labradorocoleus carpenteri, sp. nov. 
Figures 1 and 2 
Epipleural space on the base with four rows of cells, with three 
rows in the basal third, one row in the apical third. Apical third 
with A4 curved forward. Cells of elytron broader than the veins. 
Intermediate longitudinal veins for the most part straight. 
Length of elytron: 7 mm. 
Holotype: No. 87273 a,b, Invertebrate Paleontology collections, 
Princeton University. It is a well-preserved, isolated, right elytron. 
The humeral part, however, is damaged, and the bases of all the 
veins, except the anal, are impossible to distinguish. The fossil was 
collected by Professor Erling Dorf in 1958, at Redmond No. 1 
deposit in the Knob Lake District (near Schefferville) , Labrador, 
Canada. 
The species is named in honor of Professor F. M. Carpenter, 
whose courtesy and aid made possible this study. 
The discovery of this beetle, with its many tshekardocoleid-like 
features, is very interesting. Such features were lost by all other 
beetles during Upper Permian time. The labradorocoleids appear 
to have developed the complete elytral venation independently of the 
tshekardocoleids and at a later time; yet this family is not inter- 
ediate between the tshekardocoleids and more advanced forms like 
cupedids and permocupedids, for the rudimentary CuP, which is 
present in permocupedids, is absent in the labradorocoleids. Thus, 
the mechanical perfection of elytral venation was evolved in parallel 
with that of the permocupedid-cupedid line. In conclusion, then, 
the labradorocoleids arose from a tshekardocoleoid branch of archo- 
