A NEW FLIGHTLESS DOLICHOCTIS 
(COLEOPTERA: CARABIDAE) FROM SUMBAWA* 
By P. J. Darlington, Jr. 
Museum of Comparative Zoology 
The genus Dolichoctis , of the tribe Lebiini, includes a large num- 
ber of small Carabidae of the Oriental-Australian area. They are 
characterized by size (length often c. 5 mm or less), by details 
of the mouthparts (which need not be given here — see Darlington 
1968, 124), and almost always by presence of 2 minute punctures 
without setae on the posterior half of the third interval of each 
elytron. Most are arboreal, occurring often in under-story foliage 
of rain forest, although a few species probably occur in leaf litter 
on the ground. All previously known Dolichoctis are fully winged, 
so far as I know. The finding of a species in which wing atrophy has 
occurred is therefore of special interest. Atrophy of wings is in 
fact rare among all the Lebiini of the Indo-Australian islands: of 
160 members of the tribe found on New Guinea, only one has 
reduced wings ( Nototarus papua Darlington 1968, 186). Of course 
wing atrophy is very common among some other Carabidae in some 
other parts of the world (Darlington 1936; 1943). 
Dolichoctis pedestris n. sp. 
(Fig. 1) 
Description. With characters of genus; form as in Fig. 1 ; dark 
brown, reflexed elytral margins, legs, mouthparts, and antennal 
bases brownish-yellow, antennae darker from parts of 3rd segments; 
reticulate microsculpture heavy and isodiametric on front, distinct 
and somewhat transverse on pronotum, lighter and more transverse 
on elytra. Head 0.75 and 0.73 width prothorax; 2 strong pre- 
sumably formerly seta-bearing punctures over each eye. Prothorax 
subcordate; width/length 1.22 and 1.27; base/apex 0.92 and 1.02; 
lateral margins moderate and moderately reflexed, without setae or 
seta-bearing punctures; disc weakly convex, with virtually entire 
impressed middle line, transverse impressions obsolete, baso-lateral 
impressions weak, surface across base irregular but not distinctly 
punctate. Elytra short, wide; width elytra/prothorax 1.77 and 1.86 
(proportions approximate, because elytra slightly spread in all 
specimens) ; apices obliquely truncate with sutural angles narrowly 
* Manuscript received by the editor January 28, 1970 
387 
