REVISIONAL STUDY OF 
THE ORDER PALAEODICTYOPTERA IN 
THE UPPER CARBONIFEROUS SHALES 
OF COMMENTRY, FRANCE 
PART II 1 
By Jarmila Kukalova 2 
Charles University, Prague 
An introductory discussion of the Palaeodictyoptera found in the 
Commentry shales and of the collection in the Institut de Paleon- 
tologie in Paris, as well as an account of the background of this 
investigation, was included in the first part of these studies. The 
present part deals with the following seven families : Homoiopteridae, 
Lycocercidae, Graphiptilidae, Breyeriidae, Eugereonidae, Archaemeg- 
aptilidae and Megaptilidae. Compared with the Spilapteridae, con- 
sidered in Part I, all of these families are small, consisting of only 
a few genera, at least from the Commentry shales. However, they 
show much diversity of structure and indicate the extensive range of 
wing modifications which occurred in the Palaeodictyoptera, includ- 
ing the reduction and shortening of the hind wings. The third part 
of this study will deal with the Dictyoneuridae, which provide us 
with more information about the body structure in this order of 
insects. 
Family Homoiopteridae Handlirsch 
Homoiopteridae Handlirsch, 1906: 91; Lameere, 1917: 102; Handlirsch, 
1919: 16; Handlirsch, 1921: 133. 
Roechlingiidae Guthorl, 1934: 188; Kukalova, 1960: 1. 
Thesoneuridae Carpenter, 1944: 10. 
Scepasmidae Haupt, 1949: 42. 
Type genus: Homoioptera Brongniart, 1893. 
This family, as established by Handlirsch, included Homoioptera 
Brongniart, Graphiptiloides Handlirsch (1906), Homoeophlebia 
Handlirsch (1906), all from Commentry; and Anthracentomon 
Handlirsch (1904) from Belgium. Of these genera, Graphiptiloides 
(— Graphiptilus) is here assigned to the Graphiptilidae; Homoeo- 
Tublished with the aid of a grant from the Colies Fund of the Museum 
of Comparative Zoology at Harvard College and a Grant-in-Aid of Research 
from the Society of the Sigma Xi. This study has also been supported in 
part by grants numbered GB2038 and GB7038 (F. M. Carpenter, Principal 
Investigator) from the National Science Foundation. Part I, dealing with 
the family Spilapteridae, was published in Psyche, Vol. 76, pp. 163-215. 
Alexander Agassiz Lecturer in Zoology, Harvard University, 1969. 
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