452 
Psyche 
[December 
lirsch to a separate species, brongniarti , but was identified again as 
goldenbergi by Lameere (1917, p. 153). 
The type specimen (21-1) has been discussed many times by 
various authors but of these only Brongniart and Lameere actually 
studied the fossil. Most interpretations are highly speculative and 
not worth discussing here. The fossil shows so many structures 
which are important for the whole order that it deserves the most 
detailed study. Actually, the specimen might contribute even more 
details than I was able to work out in my limited stay of several 
weeks at the Paris Museum. The following discussion is based 
mostly on the obverse, with the exception of the abdominal ap- 
pendages, which are better preserved in the reverse. In figure 32, 
as usual, both obverse and reverse have been used. 
The type specimen shows fragments of two twisted wings. The 
broad wing on the right side is a fore wing, having a. broader sub- 
costal and r-rs areas than the narrower hind wing on the left side. 
This conclusion was reached after noting that the second specimen 
(21-2, shown in figure 35), with fore and hind wings in natural 
positions, showed the same differences. 
The body of specimen 21-1 is twisted in such a way that the 
thorax shows the dorsal side, while the abdomen shows the distal 
end in full lateral view. The head is in perfect frontal position. 
The insect apparently first rested with the beak oriented along the 
body axis ; later the head became loose and shifted 90° to the left. The 
prothoracic lobes lifted from the pronotum and overlapped so that 
they are now in lateral position. Such preservation is unusual for 
the Palaeodictyoptera and proves beyond any doubt that (1) the 
prothoracic lobes were not fused together to form a pronotal shield, 
as claimed by Sharov (1966) 5 ; and (2) that they were easily move- 
able. The legs of specimen 21-1 are extended on both sides of the 
body. All three right legs have a deep suture near the proximal end 
of the tibia, giving the impression of an extra segment fused with the 
tibia. I have observed similar sutures in Stenodictya agnita (Meu- 
nier) and Stenodictya oustaleti (Brongniart). It is of great interest 
that in Recent Ephemeroptera there is a marked bend in the same 
part of the tibia (e.g., Ecdyonurus). But the suture on the tibia 
does not seem to be present in the related orders Megasecoptera and 
Diaphanopterodea and not even in all Palaeodictyoptera. At any 
The prothoracic lobes were attached to the pronotum by short cuticular 
ridges in the center of their basal part. The perfectly preserved prothoracic 
lobes of Stenodictya will be described in Part III of this series of papers. 
