454 
Psyche 
[December 
rate, the part divided by the suture makes a single piece with the 
tibia, so that it probably had hardly any functional significance. 
The segments of the abdomen, undoubtedly because of decaying 
processes and distortion, show varying portions of the intersegmental 
membrane. The ovipositor is robust and opened widely. Between the 
ovipositor valves, a pair of sac-like cuticular structures appears to 
come from the abdomen. These have been incorrectly interpreted as 
gonapophyses by many authors. Demoulin (i960) considered them 
the evaginated cuticle of the oviduct and pointed to similar cases 
known in Recent Ephemeroptera after the laying of the eggs in paired 
large clusters (e.g., Polymitarciidae) . 
Above the end of the abdomen, twisted backwards, there are two 
structures described by Brongniart as “crochets dorsaux.” Detailed 
examination shows that they are “attached” to the cercus. These 
seem like parts of a smaller and much less heavily sclerotized ovi- 
positor and could be part of a cast cuticle. It is conceivable that the 
Palaeodictyoptera had an adult molt, in which case this “ovipositor” 
could have been part of the previously molted cuticle. 
The ovipositor of Lycocercus goldenbergi resembles, in its broad 
attachment to the 9th segment, the ovipositors of certain dragonflies, 
such as the Zygoptera and some Anisoptera, especially those adapted 
for endophytic oviposition. Moreover, it resembles the ovipositor of 
some Diaphanopterodea (Permian of Kansas and Czechoslovakia, un- 
published material) in the prolongation of the lateral margin of the 
9th tergite anteriorly. The surface of the gonapophyses does not 
show any sculpturing, whereas in Asthenohymen it is armed by ridges 
and even stout hairs directed backwards, as in the endophytic 
Hymenoptera. 
The following account is based on the type specimen and on 
specimen 21-2. 
Fore wing: length 73 mm, width 25 mm, broadest in the first 
third. Precostal strip extending to about the middle of the wing. 
Posterior margin with small convexities at the end of each area. 
Apex directed posteriorly, rounded. Rs with about 5 branches, the 
first two of them more or less richly branched ; MA arising occa- 
sionally distally from the first branch of MP. MP forked 11-19 
times. Cu dividing near the base, CuP with 3-5 posteriorly directed 
branches, often forked. Anal area moderately large with veins 
forked many times. Cross veins fine, dense, curved, sometimes 
branched and connected. Hind wing fragment: length 39 mm, width 
22 mm. 
Basic body structures: head small, with very large clypeus hav- 
