1969] 
Kukcilova — Palaeodictyoptera 
459 
It is to be emphasized that the hind wings of both Graphiptilus 
and Rhabdoptilus , though broad and therefore relatively short, do 
not show any sign of reduction. This fact together with the vena- 
tional pattern places them far from Lithoptilus , which has been 
erroneously associated with Rhabdoptilus (Demoulin, 1958). 
The fore wing is unknown. The hind wing is of broadly triangular 
shape. Anterior margin almost straight; Sc long; Rs area reduced 
in size, with 3-4 branches; MA simple, MP with several branches; 
CuA simple, CuP branched several times; several anal veins. Cross 
veins dense, fine, long, sometimes connected by anastomoses. Color 
markings usually in the form of irregularly distributed larger spots 
and smaller rounded spots. 
The family Graphiptilidae resembles Breyeriidae in several fea- 
tures, already noted above, including the wing markings. The graph- 
iptilids differ from the breyeriids in having a denser pattern of cross 
veins and in having the branches of the main veins obliquely oriented 
(not perpendicular to posterior margin, as in some breyeriids) and 
in having these branches closer together. Among the Breyeriidae, 
the less specialized genus Stobbsia recalls most Graphiptilidae and 
may turn out to belong to that family. 
Two genera are included in Commentry shales: Graphiptilus 
Brongniart, 1893; Rhabdoptilus Brongniart, 1893. 
Genus Graphiptilus Brongniart 
Graphiptilus Brongniart, 1893: 348; Handlirsch, 1906: 99; Handlirsch, 1919: 
20; Lameere, 1917: 150. 
Graphiptiloides Handlirsch, 1906: 92; Handlirsch, 19-9: 16. 
Type species: Graphiptilus heeri Brongniart, 1893 (SD Hand- 
lirsch, 1906). 
This genus was erected by Brongniart for heeri , williamsoni and 
ramondi. Rarnondi was referred by Handlirsch (1906, p. 100) to 
a new genus Spiloptilus , which has been transferred to the Spilap- 
teridae (Kukalova, 1969) ; williamsoni was assigned by Handlirsch 
(1906, p. 92) to a new genus Graphiptiloides and placed in the 
Homoiopteridae. As mentioned already by Lameere (1917, p. 1 5 1 ) 
heeri and williamsoni are actually very close and they are undoubtedly 
conspecific; for some reason Lameere believed that williamsoni was 
based on a fore wing, though the unique type has the same wing 
form as heeri , characteristic for hind wings in this family. 
A careful study of the specimens has made possible a reliable 
determination of the outline of the hind wings, their triangular 
