460 
Psyche 
[December 
form becoming obvious. For the relationship of Graphiptilus (as 
well as for most other Palaeodictyoptera) , the cross veins are most 
important. Those of Graphiptilus , being very fine and close to- 
gether, have not been observed or at any rate mentioned by previous 
authors. 
Hind wing very broad in proximal half; wing membrane usually 
with many elongate spots and small circular spots. Rs with four 
short branches; branches of main veins obliquely oriented to the 
posterior margin. Anal area reaching about midwing. Cross veins 
fine, dense, straight, regular. 
Graphiptilus differs from the related genus Rhahdoptilus Brongni- 
art by its fewer branches of MP, more obliquely oriented branching 
of main veins and in the more regular cross venation. The color 
pattern differs by the shape of the elongate spots. 
Only the type-species is known from the Commentry shales. 
Graphiptilus heeri Brongniart 
Figure 37, 38 
Graphiptilus heeri Brongniart, 1893: 349, pi. 19, fig. 13, Handlirsch, 1906: 
100, pi. 11, fig. 12; Handlirsch, 1919: 20; Lameere, 1917: 150. 
Graphiptilus 'williams oni Brongniart, 1893 : 350, pi. 19, fig. 12; Lameere, 
1917: 151. 
Graphiptiloides 'williamsoni Handlirsch, 1906:92; Handlirsch, 1919: 16. 
This species was based by Brongniart on a single specimen, 19-13, 
a hind wing without base but with a well preserved color pattern. 
As noted above, I consider the type of williamsoni (specimen 19-12) 
to be the same species. 
Hind wing: length about 60 mm, width 27 mm, broadest at the 
end of the first third of the wing; light spots of diverse length are 
present, limited by cross veins, and in addition small round light 
spots irregularly distributed over the wing. Anterior margin straight, 
anal area moderately rounded. Apical half of wing abruptly narrow- 
ing distally, apex rounded, shifted anteriorly to the axis of the 
wing. R-rs area only a little broadened ; anal area large with at 
least nine long branches reaching or extending beyond the middle 
of the wing; some cross veins in costal and subcostal area stronger 
than the others and more oblique. 
The differences between specimens 19-13 (type of heeri) and 19- 
12 (type of williamsoni) are in the presence or absence of small 
forks on CuA and CuP, in one branch more or less on MP and 
CuP and in the relative lengths of the anal areas. None of these 
seem to be at the specific level. 
