472 
Psyche 
[December 
of Palaeodictyoptera break off easily along this line. Thus, Cal- 
vertiellidae are usually found with the base of the anal area broken 
away. In Homoiopteridae the anal veins and cross veins are distinctly 
different before and behind the furrow, which thus forms some type 
of a separate basal area. The high percentage of palaeodictyopterous 
wings with similarly broken bases indicates the presence of the basal 
furrow in many families. At the present time I am not able to 
explain this structure. 
Wings unequal in length, the fore pair being longer. Hind wings 
shortened, with modified venation, different from that of the fore 
wings. Wing membrane usually dark with very small light spots 
restricted to individual cells of the reticulation. Fore wings long and 
very narrow, with the precostal area and precostal strip present ; deep 
furrow obliquely crossing the anal area and terminating on R; Sc 
terminating before apex; stems of R and M arising separately but 
touching for some distance beyond the base; MA simple, MP with 
2-4 branches; CuA usually simple, CuP usually with one branch 
only. Anal area long and narrow with pectinate and widely separated 
veins. A very dense pattern of coarse cross veins present, often with 
anastomoses and sometimes forming a network. 
The nearest relatives of the Eugereonidae appear to be the Archae- 
megaptilidae, Calvertiellidae and Protagrionidae ; more distantly re- 
lated are the Dictyoneuridae. All these affinities have at least 
partially been mentioned by previous authors (Handlirsch 1906, 
1937; Lameere 1917, 1935; Martynov 1932; Tillyard and Fraser 
1938; Laurentiaux 1953; Carpenter 1964; Kukalova 1964). There 
is also, in my opinion, a more distant relationship with another 
family that has markedly shortened hind wings — Megaptilidae. 
This family differs in having much broader and shorter fore wings, 
with a very large rs area but it recalls the Eugereonidae in having 
the branches of the main veins often arranged perpendicularly to the 
posterior margins; also in the simple MA and CuA, the branching 
of MP and in the very dense, richly anastomosing cross venation. 
The families mentioned above probably form a related group within 
the order. 
Genera included in commentry shales: Dictyoptilus Brongniart, 
1893; Peromaptera Brongniart, 1893. 
Other genera in other deposits: Eugereon Dohrn, 1866 (Lower 
Permian of Germany; V aldeania Teixeira, 1941 (Stephanian, 
Portugal) . 
