1969] 
Kukalovd — Palaeodictyoptera 
473 
Genus Peromaptera Brongniart 
Peromaptera Brongniart, 1893: 391; Handlirsch, 1906: 79; Lameere, 1917: 
160; Handlirsch, 1919: 11. 
Type species: Peromaptera filholi Brongniart, 1893. (OD) 
This genus is monotypic, based upon a single specimen with fore 
and hind wings preserved. Unfortunately, this remarkable fossil, 
showing a greatly reduced hind wing with the venation modified 
to the highest degree among Palaeodictyopera, could not be found 
in the collection in Paris since I began this study. The following 
account is based on Lameere’s revision (1917) and on photographs 
which Professor Carpenter made in 1938, when the specimen was 
in the collection. 
Peromaptera was referred by Brongniart to the Dictyoneuridae. 
Handlirsch (1906, p. 79) erected for it a separate family on the 
basis of the shortened hind wing. Lameere (1917, p. 160) considered 
this separation as unnecessary because of the obvious relationships of 
the genus to Dictyoptilus. 
From my studies of the photographs, I consider that Lameere’s 
conclusions are correct: not only fore wing venation and cross vena- 
tion but also all basal structures of the wing seem to be very similar 
in both Peromaptera and Dictyoptilus. The hind wing, on the other 
hand, is very different from that of both Eugereon and Dictyoptilus , 
being shorter and broader with a modified venation. But since we 
do not know the amount of individual and specific variation of the 
hind wing among the Eugereonidae and since the fore wing is almost 
indistinguishable from that of Eugereon , I do not consider it 
practical at this time to put Peromaptera in a separate family. 
Fore wing narrow, broadening at the middle part. MP with few 
branches, CuP forked. Cross veins dense. Hind wing much broader 
than fore wing, extending almost to 2/3 of the fore wing length, 
with broad apical part. Sc probably long; sc-r area broad; R 1 ending 
relatively far before apex; Rs originating very basally, rs area being 
very small ; M probably dividing beyond midwing, giving rise to 
simple MA and forked MP; CuA simple, CuP forked; anal area 
probably large, extending beyond midwing. Cross venation as in 
fore wings, in sc-r and r-rs areas much less dense than on the rest 
of the wing. 
P eromaptera differs from Eugereon and Dictyoptilus in the shape 
of the hind wing. From all other genera of Eugereonidae the hind 
wing differs in the extreme modification of the hind wing venation. 
Species included in Commentry shales: Peromaptera filholi 
Brongniart. 
