476 
Psyche 
[December 
p. 160), however, considered Archaemegaptilus related to Dicty- 
optilus (Eugereonidae) . 
Archaemegaptilus is known by a hind wing, which is not shortened 
and which has a venation similar to that of the fore wing of Dicty- 
optilus. In all genera of Eugereonidae, so far as we know, the hind 
wing is shortened and its venation is modified. The cross venation 
Archaemegaptilus is much coarser than in Dictyoptilus and forms 
intercalary sectors along the posterior margin, as in another related 
family, Calvertiellidae. Sc is almost as long as in Eugereonidae but 
terminates in a fork, forming thus an intermediate stage leading to 
an Sc which is short and which ends on R, as in Calvertiellidae. The 
stems of R and M are touching near the base as in Eugereonidae. 
The precostal area is very narrow and the wing venation less 
specialized than in both Eugereonidae and Calvertiellidae. In this 
way, Archaemegaptilus represents to some extent a combination of 
morphological features of both families being at the same time more 
primitive than any of them. I believe, therefore, that Handlirsch 
was probably correct in establishing a separate family for A rchaemeg- 
aptilus. Dictyoneurella perfecta Laurentiaux, 1949 (Westphalian of 
France), should be referred to the same family, in my opinion; it 
is a fore wing with a long narrow precostal area, intercalary sectors 
and a venation and cross venation very similar to those of Archae- 
megaptilus. 
The following account is based on Dictyoneurella (fore wing) 
and the completed figure of Archaemegaptilus (hind wing). 
Wings probably equal in length, hind pair broader. Venation in 
fore and hind wings alike. Wing membrane thin, with darker 
venation or dark with small light spots. Precostal area narrow, 
postcostal area completely missing. Sc more or less shortened. 
Stems of R and M close together or touching for some distance near 
base. Rs area small with few branches, MA simple, MP branched 
several times. CuA simple, CuP with several branches. Anal area 
with pectinate series of branches. Cross veins strong but not dense, 
forming a loose reticulation and convex intercalary sectors. 
The family Archaemegaptilidae is closely related to the Eugere- 
onidae, Calvertiellidae and Protagrionidae, from all of which it 
differs in its more primitive venation with independent branches 
and many-branched CuP. 
Genus included in Commentary shales: Archaemegaptilus 
Meunier, 1908. 
Genus from other locality: Dictyoneurella Laurentiaux, 1949 
(Westphalian, France). 
