1969] 
Kukalova — Palaeodictyoptera 
483 
Figure 49. Lithoptilus boulei (Meunier) ; hind wing. Holotype. 
Hind wing very shortened and broad, especially in the apical part. 
Anterior and posterior margins almost parallel. Sc long, subcostal 
area broad; Rs originating at about midwing; Rs area extremely 
small; M dividing at the level of the origin of Rs, MA simple, 
MP forking; Cu dividing within the first third of the wing length; 
CuA simple; CuP with several branches. Cross veins coarse and 
very dense, connected by numerous anastomoses, sometimes turning 
into a network. 
Lithoptilus Lameere differs from Megaptilus (in the hind wing) 
by the broad subcostal area, straight Sc, R and Rs; by M dividing 
more distally; and by the somewhat less dense cross venation. 
Species included in Commentry shales: Lithoptilus houlei 
(Meunier, 1908). 
Lithoptilus boulei (Meunier) 
Figure 49 
Archaeoptilus boulei Meunier, 1908 : 153; Meunier, 1908: 35, fig. 1; Meunier, 
1909: 131, pi. 1, fig. 1. 
Lithoptilus boulei Lameere, 1917: 157; Demoulin, 1958: 1-5. 
Anaxion boulei Handlirsch, 1919: 19, fig. 21. 
This species is monotypic, based by Meunier on a vestigial hind 
wing of a large palaeodictyopteron. Following the weak imprint of 
the veins on the matrix, I was able to work out some of the more 
proximal part of the type specimen. 
Hind wing fragment: length 66 mm, width 40 mm. Anterior 
