178 
Psyche 
[September 
All of the Bethyloidea are clearly identifiable as to family, but 
more difficult to place in present-day tribes and subfamilies. The 
bethylid genus Archaepyris appears to have some features of each 
of the major subfamilies, while Celonophamia appears annectant 
between the Epyrini and Cephalonomiini. Cretabythus is a truly 
enigmatic form, with many bethylid-like features (particularly the 
mandibles) but with an unusual wing venation suggesting the Scole- 
bythidae. Taimyrisphex is perhaps the most generalized wasp yet 
described from the Cretaceous. Although I have placed it in the 
Sphecidae, one might argue that it is a scolioid or even a prototype 
of the Pompilidae. We do know that the Sphecidae underwent 
considerable evolution before the end of the Cretaceous, for two 
specialized forms of the subfamily Pemphredoninae have now been 
described. 
In conclusion, it may be useful to list the Aculeata now known 
from the Cretaceous, bearing in mind that this list will be con- 
siderably augmented when all recently collected material has been 
described : 
SCOLIOIDEA 
Cretavidae: Cretavus 
?SCOLIOIDEA-BETHYLOIDEA 
PScolebythidae : Cretabythus 
BETHYLOIDEA 
Bethylidae : A rchaepyris 
Celonophamia 
Cleptidae: Procleptes 
Hypocleptes 
P rot amis ega 
References 
Evans, H. E. 
1963. A new family of wasps. Psyche, 70: 7-16. 
1969. Three new Cretaceous aculeate wasps (Hymenoptera) . Psyche, 
76: 251-261. 
Sharov, A. G. 
1957. First discovery of a Cretaceous stinging hymenopteron (Acu- 
leata). Dokl. Akad. Nauk., 112: 943-944. (In Russian). 
Wilson, E. O., F. M. Carpenter, and W. L. Brown, Jr. 
1967. The first Mesozoic ants, with the description of a new subfamily. 
Psyche, 74: 1-19. 
FORMICOIDEA 
Formicidae: Sphecomyrrna 
?SPHECOIDEA 
PSphecidae : A rchisphex 
Taimyrisphex 
SPHECOIDEA 
Sphecidae 
Pemphredoninae: Lisponema 
Pit to ecus 
