1973 ] 
Roth — Blattaria 
251 
The arrangement of the male genital phallomeres is shown in 
figure 1. In Pycnoscelus indicus all 3 phallomeres are well devel- 
oped and sclerotized (Figs. 7-9). The L2d is separated from L2vm; 
the obliquely more or less truncate ends of these 2 sclerotized struc- 
tures (Figs. 7, 10, 21, 24, 31) have the appearance of having been 
broken off and separated from L2vm. The outer lower curved por- 
tion of L2d is spicular (Figs. 1, 7, 10-13), and the underlying 
prepuce is densely “hairy” but otherwise not unusually shaped 
(Fig. 7, P). The curved genital hook (R2) lacks a subapical 
incision, is heavily sclerotized, somewhat truncate at the apex, the 
inner curved margin with (Figs. 8, 14-18) or without (Figs. 19-20) 
small projections. The Li is very well developed with the cleft 
turned upward and its margins heavily sclerotized (Figs. 1, 9). The 
genitalia of Pycnoscelus surinctmensis (Figs. 21-25), and Pyncoscelus 
nigra (Brunner) (Figs. 31-33) are indistinguishable from those of 
P. indicus. Habitus photographs of P. indicus and P. nigra are shown 
in figures 2 and 4. 
The shapes of the L2d and R2 (Figs. 26, 27, 29, 30) of Pycnos- 
celus semivitreus Princis (Fig. 3) differ from those structures in 
P. indicus ; P. surinamensis , and P. nigra; however, the shape of Li 
in all 4 of the above species is similar (cf. Figs. 9, 23, 28, 33 (dis- 
torted in preparation)). Two of the genital phallomeres of Pycnos- 
celus striata (Kirby) are distinguishable from those of the other 
species of the genus. Its I>2d (Fig. 34) differs in shape, lacks the 
spicular surface characteristic of the outer lower curved region and 
is only slightly separated from L2vm (cf. Fig. 31). The curved 
portion of R2 (Fig. 35) of striata is more elongate and slender than 
in semivitreus (Figs. 27, 29) and more uniform in width than in 
P. indicus (Figs. 14-20) or nigra (Fig. 32). 
Princis (1964) included Stilpnohlatta in the Pycnoscelidae and 
the genitalia of S. opaca (Walker) (Figs. 37-39) tends to support 
this conclusion, though I relegate his family to subfamily rank 
(McKittrick, 1964). Especially notable is the marked similarity in 
appearance of the Li of Stilpnohlatta (Fig. 39) with those of 
Pycnoscelus (Figs. 23, 36). The L2d (Fig. 37) of S. opaca is 
greatly reduced and irregular in outline and is widely separated from 
Fig. 1. Male genitalia (dorsal view). Top. (106 MCZ). Pycnoscelus 
indicus. Zamboanga, Philippine Islands, (det. Roth). Bottom. (70 BMNH). 
Proscratea complanata. Sao Gabriel, Rio Negro, Brazil, 27.IX.1927, J. F„ 
Zikan. (LI — first sclerite of left phallomere; L2d — dorsal sclerite of L2 ' r 
L2vm — ventromedial sclerite; M — saclike membrane above L2d; R2 — 
hooked sclerite of right phallomere). 
