1973] 
Gamboa & Alcock — Brochymena 
267 
(2) The male seizes the anterior or posterior dorsum of the female 
with his legs. Unreceptive females respond by breaking away 
from the male or by moving off with their partner clinging to 
them. 
(3) If the female is immobile, the male moves so as to face his 
mate while continuing to grasp the dorsum of her body. He 
then begins a rapid crablike walk from side to side over the 
anterior female dorsum. Receptive females gradually raise their 
abdomen in response to this activity. The male’s back and forth 
movements varied from 20-34 in number and lasted from 23-43 
seconds in four filmed courtships. 
(4) At some point the male moves off the anterior dorsum of the 
female and continues out along the side of his mate while con- 
stantly antennating her lateral surface. 
(5) The male, upon reaching the posterior of the female, sweeps 
his head under her elevated abdomen ; his antennae move rapidly 
up and down in alternating strokes touching the female’s 
abdominal venter. The male may prod and lift the body of 
unresponsive females which have not voluntarily raised their 
abdomens. 
(6) As the male’s head passes under the body of the female he 
begins a tight 180° turn that brings him into the end-to-end 
precopulatory position. Unreceptive females may dash off down 
a branch as the male executes this maneuver. 
(7) The male elevates his abdomen while sweeping the aedeagus in 
a zig-zag pattern against the venter of the female’s abdomen. 
Each sweep brings the male aedeagus progressively closer to 
the female genitalia; linkage occurs when mutual genital con- 
tact occurs. 
(8) Upon linkage, the insects’ bodies jerk violently from side to 
side for several minutes. The coupled pair may move a short 
distance during this time. 
(9) The male rapidly drums the sides of the female’s abdomen with 
his hindlegs (also abserved by Ruckes, 1938, for B. sulcata ). 
Spells of drumming are interrupted by pauses during which the 
male rests its hind legs on the dorsum of the female’s abdomen. 
This continues as long as the partners are linked. One pair of 
stinkbugs remained in copulo for at least 75 min but had sepa- 
rated when observed 45 min later. 
Discussion 
This report provides additional evidence that among the Penta- 
