THE MALE GENITALIA OF BLATTARIA. 
XI. PERISPHAERIINAE* 
By Louis M. Roth 
Pioneering Research Laboratory 
U. S. Army Natick Laboratories 
Natick, Massachusetts 01760 
In this paper I shall illustrate and characterize the male genitalia 
of 11 genera (indicated with an * in Table 1) of cockroaches which 
belong to the Perisphaeriinae. Three recent classifications which 
include these genera are shown in the table. I follow McKittrick 
(1964) in placing all of those discussed here under the Blaberidae: 
Perisphaeriinae, and Beier (1961) in using tribes to separate them. 
Rehn (1922) discussed the characteristics of Pilema, Cyrtotria , Dero- 
calymma , Zuluia , and Hostilia, and Princis (1963) devised keys to 
some South African species of the above genera, as well as those of 
Bantua , P erisphaeria, and Blepharodera. 
Materials and Methods 
All genitalia were treated with 10% KOH, dehydrated, cleared, 
and mounted in Permount. The source of each of the specimens 
examined is given using the following abbreviations: (ANSP) = 
Academy of Natural Sciences, Philadelphia; (BMNH) = British 
Museum (Natural History), London; (CUZM) = Zoological 
Museum, Copenhagen; (L) = Zoological Institute, Lund, Sweden; 
(HO) = Hope Department of Entomology, Oxford, England; 
(M'CZ) = Museum of Comparative Zoology, Harvard University, 
Cambridge, Mass. Locality data and names of specialists who identi- 
fied the specimens, if known, follow these abbreviations; most of the 
species were determined by Princis or Rehn. The number preceding 
the abbreviations refers to the number assigned the specimen and its 
corrsponding genitalia (on a slide) Which are deposited in their re- 
spective museums. 
Results and Discussion 
The normal arrangement of the phallomeres of male genitalia of 
the Perisphaeriinae, as seen in dorsal view, is illustrated in figure 1 . 
Based on male genitalia, I suggest the erection of 4 tribes for the 
* Manuscript received by the editor October 10, 1973 
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