86 
Psyche 
[March-June 
which is an integral part of the preputial membrane. The prepuce 
has no distinctive shape, other than this sclerotization. The Li is 
markedly reduced and, in most species, its sclerotization is restricted 
mainly to the region of the cleft. The tip of the genital hook (R2) 
is more lightly sclerotized than the main body of the hook, and in 
some species is a separate pointed or rounded lobe which may break 
off. Gurney (1965) has drawings of the genital hooks of Nauphoeta 
cuiereci and three species of TIenschout edema, but erred in indicating 
that they are part of the left phallomere. 
The genera listed earlier are arranged linearly according to 
Princis (1965). The genitalia are essentially so basically similar 
in the genera studied here that there is little reason to alter this 
arrangement. However, there are certain differences in structure 
of L>2d and. R2 which allow the genera to be placed into three 
tribes. 
1. Oxyhaloini ( Oxyhalod) (Figs. 29-46). Characteristically the 
upper right side of the L2d is extended into a long relatively narrow 
arm (Figs. 29, 3 2, 35, 38, 41, 44). The genital hook (R2) differs 
from all other genera of Oxyhaloinae. The rounded outer surface 
has minute setae and the apical lobe appears to arise from the dorsal 
surface as a thinly sclerotized membrane and extends from behind 
the tip of the darkly pigmented hook (Figs. 30, 33, 36, 39, 42, 45). 
2. Nauphoetini ( Griffiniella , Nauphoeta , Henschoutedenia, Leu- 
cophaea, and Jagrehnia) (Figs. 6-25, 47-139). The curved genital 
Figs. 2-5. Male subgenital plates of Oxyhaloinae (ventral views). 2. 
Nauphoeta cinerea. 3. Leucophaea maderae (Fab.). 4. (188 USNM) 
Henschoutedenia sordida (Shelford) (Allotype of Nauphoeta pro c era Rehn). 
Mt. Coffee, Liberia. 5. Gromphadorhina brunneri Butler. Madagascar, 
(scale — 0.5 mm). 
