1971] 
Roth — Blattaria 
1 8 1 
tribes . . .” within the Epilamprinae. However, she tentatively 
divided the 13 genera of Epilamprinae which she studied into 5 
tribes. She included Epilampra , Litopeltis, and Poroblatta (with a 
query) in the Epilamprini. 
I have found that the male genitalia of many genera of Epilam- 
prinae may be used to make tribal designations. In the present study 
the male genitalia clearly fall into 3 groups based on distinct dif- 
ferences in the L2d and prepuce. 
1. Poroblattini ( Poroblatta [Fig. 1], Colapteroblatta [Fig. 2], 
Dryadoblatta [Fig. 3], Galiblatta [Fig. 4], Nauclidas [Fig. 5]). — 
In this tribe the L2d is elongated, curved, sclerotized, tapers slightly 
toward the tip, and is separated from L2vm (Figs„ 9, 12, 15, 18, 21, 
24). Apparently there is no distinctive prepuce. The R2 has a sub- 
apical incision (Figs. 10, 13, 16, 19, 22, 25) and the shapes of Li are 
all basically similar (Figs. 11, 14, 17, 20, 26). Hebard (1919) 
claimed that Poroblatta (Figs. 9-1 1) is related to Colapteroblatta 
(Figs. 12-14) but showed closer affinity to Aero poroblatta, and the 
nearest relative of Colapteroblatta was Poroblatta. I have not seen 
any males of Aero poroblatta, but the genitalia support Hebard ’s con- 
clusion regarding a close relationship between Poroblatta and Colap- 
teroblatta. According to Hebard (1926, p. 236) Galiblatta is ap- 
parently nearest Colapteroblatta. The close relationship between 
these 2 genera is seen in their genitalia but I would place Galiblatta 
closer to Dryadoblatta (cf. Figs. 21-23 and Figs. 24-26) than to 
Colapteroblatta (Figs. 12-14). The male genitalia of Galiblatta 
cribrosa differs from G. williamsi in the shape and microscopic sur- 
face of the tip of L2d (Figs. 18, 21, in Roth, 1968). 
Rehn and Hebard (1927, p. 319) not having access to males 
tentatively assigned the West Indian species Parasphaeria nigra 
Brunner to the genus Poroblatta. Later Rehn (1930, p. 58) erected 
the genus Nauclidas using P. nigra as the type genus; he stated 
that Nauclidas “. . . belongs to the assemblage which also comprises 
Colapteroblatta , Poroblatta, Aero poroblatta , and Galiblatta .” Rehn 
placed Nauclidas nearer Galiblatta than to any of the other genera. 
The male genitalia of Nauclidas (Figs. 15-17) confirm this close 
relationship to members of the Poroblattini. 
Rehn (1930, p. 56-58) based the genus Dryadoblatta on Homalop- 
teryx scotti Shelford. He believed that Dryadoblatta was “. . . prob- 
ably as near to Pinaconota Saussure as to any other genus known at 
this writing ... In the present incomplete state of our knowledge 
of the diagnostic features of the genera placed in the Epilamprinae, 
and in the absence of any phylogenetic concept of their classification, 
it seems best to compare Dryadoblatta with Pinaconota. Future 
