1971] 
Roth — Blattaria 
191 
most Epilampra (Roth, 1971) to place them in Epilamprini; L2d 
is a variably shaped dark sclerite separated from L2vm and the 
prepuce is usually a distinctively shaped lobe covered by microtrichia 
(Fig. 39). The Ri’s of Litopeltis (Figs. 37, 40, 43, 46), and 
Cariacasia (Fig. 49) have a subapical incision and the shapes of Li 
(Figs. 38, 41, 44, 47, 50) are similar. The differences in the 
genitalia of the 3 species of Litopeltis are so minor (Figs. 36 - 47 ) 
that it would be impossible to use them to distinguish species. 
When Hebard (1920, p. 140) described the genus Litopeltis he 
stated that it . . belongs to the second section of the Perisphaerinae, 
containing Stenopilma Sauss. [•= Cyrtotria \ and its allies. To this 
section also belong the American genera Colapteroblatta , Poroblattct 
and Acroporoblatta Hebard and Mioblatta .... nearest relationship 
with Colapteroblatta exists, this indicated by the general similarity 
of tegminal and wing form and venation and limb armament.” The 
genitalia of Litopeltis (Figs. 36-47) are sufficiently different from 
those of Colapteroblatta (Figs. 12-14) to place them in different 
tribes. 
Rehn (1928, p. 190) in discussing the genus Cariacasia placed it 
in the Perisphaeriinae and claimed it was related to Litopeltis and 
Mioblatta Saussure. However, he also stated that . . the male of 
Litopeltis superficially looks more like the epilamproid genus Leuro- 
lestes [= Phoetalia ]. The relationship of the two genera here 
treated is, however, more intimate than a casual glance, even at 
individuals of the same sex, would indicate.” Phoetalia has male 
genitalia characteristic of Blaberinae and I recently assigned it to 
this subfamily (Roth, 1970b). Because of differences in tarsal arm- 
ament, Rehn (1930, p. 59) removed Litopeltis and Cariacasia 
“. . . from the vicinity of the Poroblatta complex, although their 
general appearance much suggests the latter assemblage.” The geni- 
talia of Litopeltis (Figs. 36-47) and Cariacasia (Figs. 48-50) are 
very similar showing a close relationship, and differ from those of 
Poroblattini, thus supporting Rehn’s conclusions. 
Summary 
Based on male genitalia, 8 genera of Epilamprinae are placed into 
3 tribes as follows: 
1. Poroblattini. — Poroblatta , Nauclidas , Galiblatta ) Dryado- 
blatta , and Colapteroblatta. 
2. Notolamprini. — Notolampra. 
3. Epilamprini. — Litopeltis , Cariacasia. 
