28 o 
Psyche 
[December 
The beak was not directed ventrally from the head but anteriorly 
and ventrally. 
The section of the beak in the new specimen of dcibasinskasi is 
about 3 mm. in front of the anterior margin of the head (figures 7 
and 8) ; it presumably shows the structure of the beak close to the 
head, not at a point further along the beak. As shown in the photo- 
graph (figure 9), there is a vague circular area of discoloration which 
appears to mark off the area of the beak. Within this can be seen 
sections of two pairs of stylets; these are very dark and distinct. 
The anterior pair, which are somewhat the larger, are presumably 
the mandibles, and the other pair, the maxillae. These four stylets 
are symmetrically arranged within the beak and with respect to 
each other. However, in addition to these there is a fifth stylet, 
midway between the maxillae and slightly more posterior. The sym- 
metry of the pattern formed by these five structures, as seen in sec- 
tion, is very striking. It is virtually certain that the fifth stylet is 
one of the mouthparts, presumably derived from the hypopharynx; 
the latter, of course, is a median structure and it does function as 
one of the stylets of the haustellate mouth-parts of some Diptera 
and Hemiptera. The labium, which as noted above is broader and 
thicker than the stylets in the Palaeodictyoptera, is missing from the 
section, as are the large maxillary palpi. In any event, it now seems 
clear that in the spilapterid Palaeodictyoptera, at least, there were 
five stylets in the beaks, with an arrangement which closely parallels 
that in some Recent insects. 
Genus Spilaptera Brongniart 
In the Herdina collection there are two spilapterids that appear 
to belong to the same species. Only two representatives of this 
family have been previously reported from the Francis Creek 
Shale, or, in fact, from all Pennsylvanian deposits in North America: 
Homaloneura dabasinskasi Carpenter and Mcluckiepteron luciae 
Richardson. The new specimens belong to an undescribed species, 
which we are assigning to Spilaptera , otherwise known only from the 
Commentry shales of France. 
Spilaptera differs from Homaloneura in having several long, sig- 
moidal and oblique cross veins between Ri and Rs, beyond mid- 
wing. In Homaloneura these cross veins are straight or nearly so 
and they are transverse, or only slightly oblique. CuP is usually 
unbranched in Homaloneura , although it may be forked distally, 
very near its termination. In Spilaptera CuP ranges from unbranched 
to deeply forked. The new species described below has the sigmoidal 
