298 
Psyche 
[December 
refers to the number assigned the specimen and its corresponding 
genitalia (on a slide) which are deposited in their respective museums. 
Results and Discussion 
The most characteristic feature of the male genitalia of most 
Panchlorinae is the absence of the genital hook and light pigmenta- 
tion and marked reduction of the remaining 2 phallomeres when they 
are present. McKittrick (1964, p. 72) noted the marked reduction 
in the genitalia of Panchlora nivea and pointed out that the only 
sclerotized structures are the cleft region of Li and the lightly pig- 
mented median sclerite (L2vm). However, a study of several species 
of Panchlora shows that some forms have all three phallomeres 
(Figs. 1-4) ; males that lack certain phallomeres probably evolved 
from species that had them. Based upon the presence or absence of 
genital phallomeres, the Panchlora studied here can be divided into 
the following Species Groups: 
Group 1 ( Panchlora vosseleri) . Though lightly sclerotized, all 
three basic phallomeres are present, including an L2d (Figs. 1-4). 
Group 2 (Panchlora stanleyana) . L2d is absent; Li, L2vm, 
and R2 are present (Figs. 5-7). 
Group 3 (Panchlora nivea , P. thalassina) . L2d and R2 are 
absent; Li, and L2vm, are present (Figs. 8-15). 
Group 4 (Panchlora bidentula , P. minor , P. sagax, P. dumicola, 
P. peruana). L2d, R2, and apparently L2vm are absent; only Li is 
present (Figs. 16-20). 
Group 5 (Panchlora exoleta Burmeister). Apparently no sclero- 
tized structures are present. 
Groups 1 and 2 are African and 3 to 5 are found in Central and/ 
or South America. 
In Groups 4 and 5 it is quite possible that L2vm was so lightly 
sclerotized and Li so small, and poorly defined, that they were lost 
in preparation of the genitalia. 
The male phallomeres L2vm of Achroblatta (Figs. 24-25), Biolleya 
(Figs. 21-23), and Anchoblatta (Figs. 26-29) are very lightly pig- 
mented. Li is also reduced as in most Panchlora , and L2d and R2 
are absent. 
The marked reduction in the male genital phallomeres of this sub- 
family may be related, in some way, to the mating behavior of these 
genera. In most species of cockroaches the receptive females respond 
to male courtship by palpating his dorsum, which in many genera 
possesses specialized glands which persumably produce a secretion at- 
